Balligratus humerangulus, Taxa Only, Exact Match: 2 Treatments

Balligratus humerangulus		sp. nov.	Moret, Pierre & Ortuño, Vicente M., 2017, Balligratus, new genus of wingless ground beetles from equatorial Andean montane forest (Coleoptera: Carabidae: Lachnophorini), Zootaxa 4258 (2), pp. 101-120 : 104-108	104-108
Balligratus humerangulus			Moret, Pierre & Allegro, Gianni, 2025, Illustrated checklist of the carabid beetles (Coleoptera, Carabidae) of the Otonga Nature Reserve in biogeographic comments NW Ecuador, with ecological and, Faunitaxys (New York, N. Y.) 13 (17), pp. 1-21 : 12	12