Teyujagua paradoxa, PINHEIRO ET AL., 2016
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publication ID |
https://doi.org/10.1093/zoolinnean/zlz093 |
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persistent identifier |
https://treatment.plazi.org/id/03EB87BE-8C50-FFC2-FCA9-FB2FFDBB504C |
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treatment provided by |
Plazi |
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scientific name |
Teyujagua paradoxa |
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TEYUJAGUA PARADOXA PINHEIRO ET AL., 2016
Holotype: UNIPAMPA 653, the holotype and, so far, only known specimen of Teyujagua paradoxa consists of an almost complete, well-preserved skull articulated with the complete lower jaws, the atlas–axis complex, cervical vertebrae III and IV and some tiny fragments of cervical vertebra V ( Figs 1 View Figure 1 , 2 View Figure 2 ; Table 1).
Type horizon and locality: UNIPAMPA 653 was recovered from a fine sandstone layer with abundant carbonaceous concretions, about 5 m from the base of ‘outcrop 5’, Bica São Tomé locality ( Da-Rosa et al., 2009), Lower Triassic Sanga do Cabral Formation ( SCF), Brazil (29°36 ′ 56 ″ S, 55°03 ′ 10 ″ W). The outcrop is dominated by fine reddish sandstones intercalated with coarse sandstones and intraformational conglomerates, indicating a vast alluvial plain occasionally flooded by shallow, braided streams ( Zerfass et al., 2003; Da-Rosa et al., 2009; Pinheiro et al., 2016; Dias-da-Silva et al., 2017). An Induan– Olenekian age is inferred for SCF based on the presence of the parareptile Procolophon trigoniceps Owen, 1876 , allowing the correlation between SCF and the upper Katberg Formation of the South African Karoo Basin ( Botha & Smith, 2006; Dias-da-Silva et al., 2006, 2017). The type locality of Teyujagua paradoxa has already yielded the capitosauroid temnospondyl Tomeia witecki Eltink et al., 2016 ( Eltink et al., 2017), still undescribed archosauromorph remains and abundant cranial and postcranial procolophonoid bones, including fairly complete skulls of P. trigoniceps ( Da-Rosa et al., 2009; Dias-da-Silva et al., 2017; Silva-Neves et al., 2018). Tanystropheid archosauromorphs were also reported for other classic SCF localities ( Oliveira et al., 2018).
Emended diagnosis: Teyujagua paradoxa differs from all other known archosauromorphs on the basis of the following unique combination of characters (autapomorphies indicated by *): large, confluent external nares; external antorbital fenestrae absent; open lower temporal bars; lateral mandibular fenestrae present and positioned beneath the orbits when the lower jaw is occluded*; premaxillae lack anterodorsal processes; premaxillae bear posterodorsally directed palatal processes; anterior maxillary foramina absent; medial antorbital fossae present in maxillae; nasals are completely dorsal elements; lacrimals are broad and fill the space between the ascending and posterior processes of the maxillae; frontals have a small contribution to orbital rims; posterolateral processes of parietals elevated well above the skull roof; dorsal borders of the supratemporal fenestrae level with the dorsal margins of the orbits; squamosals with elongate ventral processes that reach a point level with the ventral margins of the orbits; wide, anteriorly open quadrate foramen; triangular supraoccipital; splenials exposed in lateral view; surangular shelves present; labiolingually compressed marginal teeth; marginal teeth distally carinated and bearing serrations; pterygoid dentition with a single tooth row on zone T 3, zone T 2 with two rows and zone T 4 present*; strong longitudinal lamina on the lateral surface of the axial centrum*; neural spine of cervical vertebra III with a rounded, posterior projection*.
COMPARATIVE DESCRIPTION
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Royal British Columbia Museum - Herbarium |
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Tavera, Department of Geology and Geophysics |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.