Roxasella umbella, Lu & Dietrich & Webb & Zhang, 2025

Lu, Lin, Dietrich, Christopher H., Webb, M. D. & Zhang, Yalin, 2025, Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species, Zoological Journal of the Linnean Society 204 (3) : -

Roxasella umbella sp.nov.

( Fig. 18A–Q) (URN:LSID:ZOOBANK.ORG:ACT: F312D0B1-CABB-4397-83D6-902881CDA95F)

Diagnosis: Head with orange markings. Crown with narrow black band at anterior margin. Male pygofer with caudodorsal margin narrowly rounded. Style apex broad and falcate. Connective short. Aedeagus with two pairs of long apical processes nearly straight distally. Without additional sclerite between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 12 setae, AV with five stout setae.

Description: Length (including tegmen): male 8.0 mm.

Coloration: Body dull stramineous. Crown with narrow black band at anterior margin and broader ochraceous band between anterior corners of eyes. Face dull stramineous dorsally, becoming paler ventrally, with narrow black band at dorsal margin. Pronotum mostly greenish brown. Exposed mesonotum stramineous except basolateral triangles greenish brown.

Crown parabolic with anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 12 setae, AV with five stout setae.

Male genitalia: Pygofer with caudodorsal margin narrowly rounded. Style apophysis slightly expanded, apex broad and falcate, without distinct posteromedial lobe, pre-apical lobe very weakly developed and obtuse. Connective short, arms slightly longer than stem; aedeagus with dorsal apodeme slightly broadened near apex in posterior view; shaf moderately long and narrow, in lateral view curved dorsad at base then relatively straight, broader at base and apex than near middle, in posterior view narrow and nearly parallel sided, with two pairs of long apical processes extended basolaterad to midlength of shaf, one pair extended slightly posterad, other pair extended slightly anterad of shaf; gonopore apical.

Material examined: Holotype: ♂, Malaysia, Sabah, FMU10, Apin apin, Keningau (05°44.31 ′ 07N, 116°45.15 ′ 72E), 1217 m, 24 April–03 May 2016, coll. Z.S. Song ( NWAFU).

Etymology: Te specific epithet refers to the umbel-like apex of the aedeagus in ventral view.

Remarks: Tis species closely resembles R. muluensis in form and coloration but differs in having the aedeagal shaf relatively straight in lateral view with the processes more slender, the stem of the connective shorter and narrower, and the crown with anterior and posterior margins parallel.