Pseudoliogenys bidentula ( Fairmaire, 1892 )

Smith, Andrew B. T. & Ruiz-Manzanos, Eider, 2010, Taxonomy and Classification of the South American GenusPseudoliogenysMoser (Coleoptera: Scarabaeidae: Melolonthinae: Diplotaxini), The Coleopterists Bulletin 64 (3), pp. 272-276 : 274-275

Pseudoliogenys bidentula ( Fairmaire, 1892) ( Figs. 1–4 )

Liogenys bidentulus Fairmaire 1892: 243 Pseudoliogenys flavidus Moser 1919: 359

( new synonymy)

Liogenys bidentula Fairmaire ; Blackwelder 1944: 227

Pseudoliogenys flavida Moser ; Blackwelder 1944: 227

Original Combination. Liogenys bidentula Fairmaire, 1892: 243 (emended from the original spelling of Liogenys bidentulus to match the feminine gender of the genus).

Type Locality. “ Rioja ”.

Type Specimen. Lectotype male at the Muséum National d’ Histoire Naturelle, Paris labeled a)

“ MUSÉUM PARIS / 1906 / Coll. Léon FAIRMAIRE” (typeset), b) “ TYPUS ” (red label, typeset), c) “ Liogenys / bidentulus / Fairm. / Rioja / [illegible word]” (handwritten), d) “ LIOGENYS / BIDENTULA / FAIRMAIRE, 1892 / LECTOTYPE / A.B. T. SMITH ♂ ” (red label, handwritten and typeset), e) “ PSEUDOLIOGENYS / BIDENTULA / (FAIRMAIRE) ♂ / Det:A.B. T.Smith 2009” (handwritten and typeset). Lectotype here designated. A lectotype is designated for this species name to preserve the stability of nomenclature by selecting one specimen as the sole, name-bearing type of the taxon.

Synonym. Pseudoliogenys flavida Moser, 1919: 359 (emended from the original spelling of Pseudoliogenys flavidus to match the feminine gender of the genus).

Type Locality. “ Argentinien ( Mendoza)”.

Type Specimen. Lectotype male at the Museum für Naturkunde der Humboldt-Univertität, Berlin labeled a) “ Mendoza / Argentinien ” (handwritten and typeset), b) “ Pseudoliogenys / flavidus Mos / Type ♂ ” (handwritten), c) “ TYPE ” (red label, typeset), d) “ SYNTYPUS / Pseudoliogenys flavidus Moser 1919 / labelled by MNHUB 2005 ” (red label, typeset), e) “ PSEUDOLIOGENYS ♂ / FLAVIDA MOSER / 1919 / LECTOTYPE / E. RUIZ- MANZANOS” (red label, handwritten and typeset), f) “ PSEUDOLIOGENYS ♂ / BIDENTULA (FAIRMAIRE) / E. Ruiz-Manzanos Det. 2010” (handwritten and typeset) . Lectotype here designated. A lectotype is designated for this species name to preserve the stability of nomenclature by selecting one specimen as the sole, name-bearing type of the taxon.

Three paralectotypes at the Museum für Naturkunde der Humboldt-Univertität, Berlin: 1 female labeled a) “ Argentinien / Mendoza ” (handwritten and typeset), b) “ Pseudoliogenys / flavidus Mos / Type ♀ ”(handwritten), c) “ TYPE ”, d) “ SYNTYPUS / Pseudoliogenys / flavidus Moser, 1919 / labelled by MNHUB 2005” (red label, typeset), e) “PSEUDO- LIOGENYS ♀ / FLAVIDA / MOSER, 1919 / PARALECTOTYPE / E. RUIZ-MANZANOS” (yellow label, handwritten and typeset), f) “PSEUDO- LIOGENYS ♀ / BIDENTULA (FAIRMAIRE) / E. Ruiz-Manzanos Det.2010” (handwritten and typeset); 1 male labeled a) “ Mendoza / Argentinien ” (handwritten and typeset), b) “ SYNTYPUS / Pseudoliogenys / flavidus Moser, 1919 / labelled by MNHUB 2005” (red label, typeset), c) “ PSEUDOLIOGENYS ♂ / FLAVIDA / MOSER, 1919 / PARALECTOTYPE / E. RUIZ-MANZANOS” (yellow label, handwritten and typeset), d) “ PSEUDOLIOGENYS ♂ / BIDENTULA (FAIRMAIRE) / E. Ruiz-Manzanos Det. 2010” (handwritten and typeset); 1 male labeled a) “ Argentinien / Mendoza ” (handwritten and typeset), b) “ Pseudoliogenys / flavidus Mos ” (handwrit-

ten), c) “ SYNTYPUS / Pseudoliogenys / flavidus Moser, 1919 / labelled by MNHUB 2005” (red label, typeset), d) “ PSEUDOLIOGENYS ♂ / FLAVIDA / MOSER, 1919 / PARALECTOTYPE / E. RUIZ- MANZANOS” (yellow label, handwritten and typeset), e) “ PSEUDOLIOGENYS ♂ / BIDENTULA (FAIRMAIRE) / E. Ruiz-Manzanos Det.2010” (handwritten and typeset).

Description. Length: 7.9–11.2 mm. Color: reddish tan head, pronotum; tan elytra. Head: clypeus deflexed at least 45° compared with base of head, clypeus narrow with 2 well-defined teeth at apex; frons with curved ridge extending between eyes, rugose, setose with long setae extending toward base of head; mouthparts greatly reduced except for maxillary palps; mentum with long, dense setae. Pronotum: widest at middle; disc with sparsely scattered, large punctures; apex with distinct border, border thicker medially. Elytra: striae weakly defined. Pygidium: almost circular in shape, completely visible beyond elytral apex; surface flat in males, inflated laterally in females. Legs: procoxae conical, protibiae with three lateral teeth; mesocoxae contiguous; tarsomere 1 distinctly longer than tarsomere 2, except in metatarsus; metafemora widened, swollen; metatibiae expanded towards apex and with a vestigious and a complete strong carina at middle; claws cleft in males and simple in females. Parameres: apex rounded, not inflated; without grooves or modified structures.

Specimens Examined. One hundred and ten specimens were examined from: Andrew B. T. Smith Collection, Ottawa, ON, Canada; Canadian Museum of Nature, Ottawa, Canada ( B. Anderson, F. Génier); Canadian National Collection of Insects, Ottawa, ON, Canada ( P. Bouchard); Fundación e Instituto Miguel Lillo, Universidad Nacional de Tucumán, Tucumán, Argentina (M. V. Colomo); Henry and Anne Howden Collection, Ottawa, Canada (now deposited at Canadian Museum of Nature ); Instituto Argentino de Investigaciones de las Zonas Áridas, Mendoza, Argentina ( F. C. Ocampo); Lyman Entomological Museum, McGill University, Ste. Anne de Bellevue , PQ, Canada ( T. Wheeler, S. Boucher); Museo Argentino de Ciencias Naturales , Buenos Aires, Argentina ( A. Roig); Museo de La Plata , La Plata, Argentina ( A. Lanteri); Museo Nacional de Historia Natural , Santiago, Chile ( M. Elgueta); Museum für Naturkunde der Humboldt-Univertität, Berlin, Germany ( J. Frisch, J. Willers); Muséum National d’ Histoire Naturelle, Paris, France ( O. Montreuil); The Natural History Museum, London, U.K. ( M. Barclay); Naturhistorisches Museum, Basel, Switzerland ( E. Sprecher) .

Distribution ( Fig. 5 ). ARGENTINA (103): Catamarca (1): 6 km N de Belen; Córdoba (3): Anisacate, Río Cuarto; La Rioja (24): La Rioja, Patquía, provincial record only; Mendoza (36):

Figures

Figs. 1–2. Pseudoliogenys bidentula male. 1) Dorsal view; 2) Oblique view.

Figs. 3–4. Pseudoliogenys bidentula female. 3) Dorsal view; 4) Oblique view.

Fig. 5. Distribution of Pseudoliogenys bidentula.