Onthophagus landolti, Harold, 1880

Arriaga-Jiménez, Alfonsina, Escobar-Hernández, Fernando, Rös, Matthias & Kohlmann, Bert, 2020, The establishment of the Onthophagus anthracinus (Coleoptera: Scarabaeidae) species complex and the description of a new species, The Canadian Entomologist 152 (1), pp. 1-17 : 14

The Onthophagus landolti species group

Boucomont (1932) published a synopsis of the Neotropical Onthophagus . In this work, he established what he called the “8 th group” of Onthophagus , encompassing the species of what is now called the O. landolti species group: O. anthracinus Harold, 1873 , O. arizonensis Schaeffer, 1909 , O. atrosericeus Boucomont, 1932 , O. chryses Bates, 1887 , O. columbianus Boucomont, 1932 , O. cribricollis Horn, 1881 , O. igualensis Bates, 1887 , O. iodiellus Bates, 1887 , O. hoepfneri Harold, 1869 , O. landolti Harold, 1880 , O. lecontei Harold, 1871 , O. longimanus Bates, 1887 , and O. rufescens Bates, 1887 .

Later, Zunino and Halffter (1997) redefined this group and renamed it the O. landolti species group and added: O. aciculatulus Blatcheley, 1928 , O. alluvius Howden and Cartwright, 1963 , O. brachypterus Zunino and Halffter, 1997 , O. digitifer Boucomont, 1932 , O. knausi Brown, 1927 , O. knulli Howden and Cartwright, 1963 , O. lebasi Boucomont, 1932 , O. mariozuninoi Delgado, Navarrte and Blackaller, 1993 , O. mextexus Howden and Cartwright, 1970 , O. schaefferi Howden and Cartwright, 1963 , O. subopacus Robinson , O. tuberculifrons Harold, 1871 , and O. zapotecus Zunino and Halffter, 1988 . Subsequently, Howden and Génier (2004) incorporated the following species to the O. landolti species group: O. altivagans Howden and Génier, 2004 , O. canelasensis Howden and Génier, 2004 , O. dubitabilis Howden and Génier, 2004 , O. gibsoni Howden and Génier, 2004 , and O. pedester Howden and Génier, 2004 and established the O. lecontei – O. subopacus species complex.

Recently, Moctezuma et al. (2016) described another species for the O. landolti species group, O. martinpierai Moctezuma, Rossini, Zunino, Halffter, 2016 . They also established another species complex consisting of O. martinpierai , O. mariozuninoi , and O. dubitabilis .

We present below an updated list of the species belonging to the O. landolti species group, including three species complexes, the O. lecontei – O. subopacus species complex as defined by Howden and Génier (2004), the O. mariozuninoi species complex as defined by Moctezuma et al. (2016), and the O. anthracinus species complex as defined in this paper. Onthophagus columbianus Boucomont, 1932 drops from this list because it was recently placed in junior synonymy with Onthophagus elegans Klug, 1832 , a Madagascan species ( Rossini and Vaz-de-Mello 2016).