Ochlata, Zhang & Cong & Shen & Song & Grishin, 2025

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2025, Advancing butterfly systematics through genomic analysis, The Taxonomic Report of the International Lepidoptera Survey 12 (5), pp. 1-201 : 167

Ochlata Grishin , new subgenus

http://zoobank.org/ 6E20DE25-20C7-4367-8A95-0FE0044C2C88

Type species. Hesperia venata Bremer & Grey, 1853 .

Definition. In the genomic phylogeny of Ochlodes Scudder, 1872 , the last prominent clade consists of all Old World species and represents a new subgenus ( Fig. 131). This new subgenus differs from its relatives by a combination of the following characters: the two parts of the stigma are at least slightly offset from each other; ventral hindwing and the apex of ventral forewing are yellow to brown, sometimes with olive overscaling but not bright-green; valva may have a prominent cleft between harpe and ampulla in some species, while in others harpe touches the ampulla, the aedeagus is narrower, with a long style and several cornuti. In DNA, a combination of the following characters is diagnostic in the nuclear genome: aly2682. 1.6:G81A, aly2178.36.1:C129T, aly 2250.11.1:A174G, aly256.7.8: T177 A, aly159.16.1:G96A; and in COI barcode: A85T, 232T or C (not A), T325 A, T514 T, T520 T .

Etymology. The name is a fusion of the names of the genus and the type species of the subgenus: Ochl [odes] + [ven] ata. The name is a feminine noun in the nominative singular.

Species included. All Old World species of Ochlodes Scudder, 1872 : the type species (i.e., Hesperia venata Bremer & Grey, 1853 ), Pamphila bouddha Mabille, 1876 , Pamphila brahma Moore, 1878 , Augiades crataeis Leech, 1893 , Ochlodes flavomaculata Evans, 1949 , Augiades formosana Bremer & Grey, 1853 , Ochlodes hasegawai Chiba & Tsukiyama, 1996 , Ochlodes klapperichii Evans, 1940 , Ochlodes lanta Evans, 1939 , Ochlodes linga Evans, 1939 , Pamphila ochracea Bremer, 1861 , Ochlodes sagitta Hemming, 1934 , Augiades similis Leech, 1893 , Pamphila siva Moore, 1878 , Hesperia subhyalina Bremer & Grey, 1853 , Papilio sylvanus Esper, 1777 , Pamphila thibetana Oberthür, 1886 , including taxa currently treated as their subspecies and synonyms.

Parent taxon. Genus Ochlodes Scudder, 1872 .