Megalophaedusa (Nordsieck, 1998)

Motochin, Risho, Wang, Min & Ueshima, Rei, 2017, Molecular phylogeny, frequent parallel evolution and new system of Japanese clausiliid land snails (Gastropoda: Stylommatophora), Zoological Journal of the Linnean Society 181, pp. 795-845 : 832

Megalophaedusa ( sensu lato)

The following Megalophaedusa species that cannot be assigned to the subgenera listed above are provisionally placed in Megalophaedusa ( sensu lato): M. aenea (Pilsbry, 1903) ; M. ikenoi (Minato, 1980) ; M. pachyspira (Pilsbry, 1902) ; M. rex ( Pilsbry, 1905) .The taxonomic status of these species must be revised after their phylogenetic relationships within the genus have been resolved definitely.

GENUS TOSAPHAEDUSA EHRMANN, 1929 (= CLADE C)

Type species: Clausilia cincticollis Ehrmann, 1900

Diagnosis: Although, Tosaphaedusa redefined here cannot be diagnosed by morphology, its monophyly is strongly supported by molecular phylogeny. The presence of unique deletions and substitutions in the hyper variable region of ITS2 ( Fig. 5) is a molecular diagnostic character of the genus. The distribution is endemic to Shikoku.

Morphology: Shell small to medium-sized. Apex rounded or somewhat pointed. Plicae and clausilium located ventrally or laterally. Superior lamella developed. Inferior lamella receded. Lunella present or absent. Principal plica developed. Upper and lower palatal plicae present. Middle palatal plica(e) present or absent. Dorsal crest present or absent. Clausilium wide or narrow. Diverticulum of bursa copulatrix developed.