Miturga australiensis (L. Koch, 1873 ), 2015

Raven, Robert J., 2015, & lt; strong & gt; A revision of ant-mimicking spiders of the family Corinnidae (Araneae) in the Western Pacific & lt; / strong & gt;, Zootaxa 3958 (1), pp. 1-258 : 23

Miturga australiensis (L. Koch, 1873) , new combination

Liocranum australiensis L. Koch, 1873: 429 .

Medmassa australiensis : Simon, 1898: 195.

Remarks. Although L. Koch (1873: 429) refers to the description in L. Koch (1867: “366”), Bonnet (1957: 2747) and others found that the last page in present copies of L. Koch (1867) is 352. Hence, the valid date should be that of L. Koch (1873) who however states that he no longer has the specimen and merely repeats the description from L. Koch (1867).

Simon (1898) suggested (“peut-être”) that the species should be included in Medmassa . No type specimen is known, but the description refers to a spider with a carapace length of 4.75 mm (large for a corinnid), with leg IV only slightly longer than I (12 vs 10.5 mm). Also considering the eyes (similar sizes and spaced with wider back row), spination (no strong pairs), colour (black spots on dorsal abdomen) and presumed claw tufts; it is most likely a species of Miturga to which the species is here transferred, if in fact it is not a nomen nudum.

TAXONOMY

Key to corinnid subfamilies (and similar genera) in the Pacific

1. Posterior eye row clearly recurved; carapace long with median transverse saddle, resembling model ant thorax ( Fig. 14f )............................................................................ Castianeirinae (part) Melanesotypus Posterior eye row strongly procurved to slightly recurved...................................................... 2

2(1). Posterior eye row clearly procurved (e.g., Fig. 16b–f )......................................................... 3 Posterior eye row recurved or almost straight ( Fig. 16a ).............................. Castianeirinae , part ( Disnyssus )

3(2). Abdomen strongly sclerotised in both sexes and strong paired spines on tibiae and metatarsi I, II ( Deeleman-Reinhold, 2001, fig. 8 as in Medmassa christae , Fig. 11d ); maxillae with distinct transverse groove ( Fig. 5b, d )........................ 7 Not as above......................................................................................... 4

4(3). Sternum with anterior ledge forming depressions at each anterior "shoulder" ( Fig. 9b ); male palpal bulb oval, with tegular apophyses ( Fig. 10a )...................................................................... Corinninae , Creugas Sternum without anterior depressions ( Fig. 5a ); male palpal bulb pyriform/pear-shaped, without tegular apophyses but may have complex embolus tip (e.g., Fig. 95a ).................................................................. 5

5(4). Very strong paired spines on tibiae and metatarsi I, II ( Fig. 11d )......................................... Medmassa Spines not strong on tibiae and metatarsi I, II................................................. Castianeirinae , part

6(3). Posterior median eyes round, domed; females with triad of spigots on PMS (as in Leichhardteus kroombit , Fig. 8b )................................................................................................... Liocranidae 7 Posterior median eyes oval, flat; females without triad of spigots on PMS............................. Phrurolithidae 8

7(6). Prolateral spine on femur I absent in males and females, and margins of the carapace smooth (see Deeleman-Reinhold, 2001, figs 348, 350)................................................................................ Oedignatha Prolateral spine on femur I present in males and females, and margins of carapace undulating (see Deeleman-Reinhold, 2001, figs 375, 386)..................................................................................... Koppe

8(6). Strong paired spines ventrally on tibiae and metatarsi I, II............................................... Orthobula No paired spines ventrally on tibiae and metatarsi I, II................................................. Sphingius

Figures

FIGURE 5. Corinnidae, female, sternum, labium, maxillae and chelicerae, ventral view, a–d: a, Battalus byrneae sp. nov.; b, d, e, Koppe sp., female (Liocranidae); c, Nyssus coloripes Walckenaer, 1805; e, f, retrolateral coxa I, showing retrocoxal hymen, f, Nyssus yuggera sp. nov.

FIGURE 8. Spinnerets, female, ventral view. Copa flavoplumosa Simon, 1886, (identified from SEM images by Charles Haddad, 2005 (Zimbabwe: Batoca Gorge & Dibu Dibu River, 17°58'S, 25°57'E, 27–28.X.1990, V. D. & B. Roth, 14 ♂ 5♀, CAS, image by M. Ramírez), (a); Leichhardteus kroombit Baehr & Raven, 2013 (b).

FIGURE 9. Creugas gulosus Thorell, 1878, female: a, habitus; b, venter; c, epigyne, ventral view; d, coxa I (right) and II (left), showing retrocoxal hymen (arrow).

FIGURE 10. Creugas gulosus Thorell, 1878, male, scanning electron micrograph, male palp, ventral views. a, cymbium and bulb; b, embolus and conductor; c, tibial apophysis, ventral view (d).

FIGURE 11. Medmassa christae sp. nov., holotype female, QM S12449: a, habitus; b, c, epigyne, external (b), and internal (c); d, femur I showing strong paired spines ventrally on tibia and metatarsus and proventral femoral spine (arrow).

FIGURE 14. Melanesotypus guadal, gen. et sp. nov., allotype female (a–d, f) & holotype male (e), BMNH: a, habitus; b, pedicel, dorsal view; c, carapace, dorsal view; d, cephalothorax and abdomen, ventral view; e, f, habitus, lateral views of male (e) and female (f).

FIGURE 16. Australian Castianeirinae, habitus: a, Disnyssus helenmirrenae sp. nov.; b, Nyssus insularis (L. Koch); c, Nyssus jaredwardeni sp. nov., d, Leichhardteus bimaculatus Baehr & Raven, 2013; e, Kolora cooloola sp. nov.; f, Ozopa colloffi sp. nov.

FIGURE 95. Ozcopa monteithi sp. nov.: a–c, scanning electron micrographs; a, b, male bulb (a) and embolic process (b), ventral view; c, paracymbial spine, ventral view; d, e, allotype female, QM S21925: epigyne, external (d), and internal (e).