Mayetia (Mayetia) ramon Novoa & Baselga, 2002

Novoa, Francisco & Baselga, Andrés, 2002, A New Mayetia Mulsant And Rey (Coleoptera: Staphylinidae: Pselaphinae) From Galicia (Northwest Spain), The Coleopterists Bulletin 56 (4), pp. 541-546 : 542

Mayetia (Mayetia) ramon Novoa & Baselga , new species

( Figs. 1–8 )

Material Examined. All specimens collected by F. Novoa and A. Baselga. Holotype male: SPAIN, A Coruña, A Capela , Monastery of Caaveiro , UTM 29TNJ7507, 80 m altitude, 11 February 1999. Allotype female: same locality, 22 January 2000. Holotype and allotype are deposited in the Museo Nacional de Ciencias Naturales, Madrid, Spain (number 12476 in the Catalogue of Type Material). Paratypes: SPAIN, A Coruña, A Capela , Monastery of Caaveiro , UTM 29TNJ7507, 80 m altitude, 6 November 1998, 3 males and 2 females; 11 February 1999, 1 male; 23 March 1999, 2 males and 2 females; 30 June 1999, 1 male and 1 female; 22 January 2000, 2 males and 4 females; 3 April 2000, 6 males and 8 females; A Coruña, Monfero , Cerqueiros , UTM 29TNJ8202, 540 m altitude, 20 May 1999, 1 male; 14 July 1999, 4 males and 4 females. Paratypes are deposited in the collection of the Departamento de Biología Animal, Universidad de Santiago de Compostela, Spain .

Etymology. The species is named for Ramón Novoa , son of the first author, in tribute to his collaboration on different expeditions to collect Coleoptera .

Diagnosis. Labrum dentate. Male metatrochanters quadrate, bearing a curved tooth. Horse­shoe shaped cleft of male pygidial sternite with right side angularly curved. Aedeagus elongate, bifurcate at apex, with the longer branch curved backwards.

Description. Length 1.1–1.4 mm. Eyeless, apterous and depigmented species. Color pale brown. Body elongate and somewhat flattened, covered with fine punctures and furnished with a clothing of soft hairs. Head: Form trapezoidal, as wide as long. Anterior border of labrum medially sinuate, with two teeth on each side, the lateral ones shorter than the medial ones ( Fig. 2 ). Antennae 11­segmented: antennomeres 1–2 enlarged; antennomeres 10–11 forming a club; antennomere 11 with a circlet of inflated setae. Both mandibles with simple tooth, never bifid ( Fig. 3 ). Maxillary palpi five­segmented, the basal segment small, second largest, third and fourth bearing a sensory development, consisting of a slightly pedunculate base with a minute swelling ( Fig. 4 ). Last segment rudimentary, much shorter and narrower than penultimate segment. Thorax: Pronotum: ovate, longer than wide, its greatest width before middle; sides slightly convergent in basal half; base arcuate. Elytra: base narrow, more or less concealed under pronotum; humeri slightly indicated; sides moderately widened posteriad; maximum width after middle; hind angles rounded and sutural angles perpendicular. Male metatrochanters quadrate, as long as broad, bearing a slightly curved tooth ( Fig. 5 ). Female metatrochanters not dentate. Tarsi two­segmented: basal joint shorter, second bearing a claw. Abdomen: males with posterior margin of pygidial sternite (sixth apparent) bearing a large horse­shoe shaped cleft ( Fig. 6 ); sternal valves of genital segment (seventh apparent) large and asymmetrical. Females without cleft in pygidial sternite and sternal valves lacking. Aedeagus elongate and asymmetrical ( Fig.7 ), composed of a short and sclerotized basal swelling, followed by a constriction; a median part irregular, more or less rotated, with a flattened expansion on right side; apex bifurcate, with the longer branch bent backwards. Female genital complex longer than wide, anteriorly with sclerotized mushroom­like armature ( Fig. 8 ).

Distribution. The new species is known only from the Natural Park of Fragas del Eume, located near the Atlantic coast in Galicia (northwest Spain).

Ecological Note. M. ramon was found in a Atlantic mixed forest composed mainly of Quercus robur Linne´, Castanea sativa Miller , Corylus avellana Linne´, and Laurus nobilis Linne´. The species occurs in leaf litter, especially in the humus layer. The ratio of females to males in the samples was approximately 1:1.

Figures

Fig. 1. Dorsal habitus of Mayetia ramon, paratype male. Scale bar: 0.25 mm.

Figs. 2–8. Mayetia ramon. 2) Labrum; 3) right mandible, ventral view; 4) maxillary palp; 5) male metatrochanter; 6) male pygidial sternite; 7) aedeagus, inside abdomen, ventral view; 8) female genital complex, ventral view. Scale bar: 0.05 mm. Figs. 9–10. Mayetia perezinigoi. 9) Male metatrochanter; 10) aedeagus, inside abdomen, ventral view. After Outerelo (1980).