Heinrichiellus vedani Ranjith, 2025

Ranjith, A. P., Broad, Gavin R., Santos, Bernardo F. & Butcher, Buntika A., 2025, First report of the genus Heinrichiellus Tereshkin, 2009 (Hymenoptera, Ichneumonidae) from the Oriental region with the description of three new species, Journal of Hymenoptera Research 98, pp. 757-778 : 757-778

publication ID

https://doi.org/10.3897/jhr.98.158760

publication LSID

lsid:zoobank.org:pub:C198839C-397F-466C-94CC-F1844CFB5F36

DOI

https://doi.org/10.5281/zenodo.16989774

persistent identifier

https://treatment.plazi.org/id/A9E07C85-1869-53BA-8355-E05F8D8AF349

treatment provided by

Journal of Hymenoptera Research by Pensoft

scientific name

Heinrichiellus vedani Ranjith
status

sp. nov.

Heinrichiellus vedani Ranjith sp. nov.

Material examined.

Holotype: • female, India: Karnataka, Chamarajanagar, Biligiri Rangaswamy Temple Tiger Reserve , scrub jungle, 12.0141.4'N, 77.0655.1'E, 4.x–17.xi.2005, Malaise trap, coll. D. R. Priyadarsanan ( ZSIK: ZSI/WGRC/I.R.-INV.29361 ) . Paratypes: • 2 females and 2 males with same data as holotype ( ZSIK) .

Holotype.

♀, body length 6.0 mm, fore wing length 4.5 mm.

Description of female.

Head. Head 1.1 × as wide as high in anterior view, 1.9 × as wide as long in dorsal view (Fig. 7 B, C View Figure 7 ); ocellar-ocular length (OOL) / postero-ocellar length (POL) = 0.7. Frons slightly convex above each antennal socket (Fig. 7 C View Figure 7 ); frons granulate without small and distinct tubercle between antennal sockets (Fig. 7 C View Figure 7 ); antennal sockets large and protruding anteriorly in dorsal view (Fig. 7 C View Figure 7 ); face 1.2 × as wide as high, minutely punctate associated with oblique striations medially forming ‘ V’ - shaped area (Fig. 7 B View Figure 7 ); inner margins of compound eyes slightly diverging ventrally (Fig. 7 B View Figure 7 ); clypeus strongly convex and transverse, 2.2 × as wide as high, with medially concave apical margin (Fig. 7 B View Figure 7 ); anterior tentorial pits large (Fig. 7 B View Figure 7 ); labrum very wide, 0.5 × as wide as clypeus, with straight apical margin, protruding from under clypeus (Fig. 7 B View Figure 7 ). Mandible slender, 2.1 × as long as wide at base, gradually narrowed from base to apex, with indistinct lower tooth (Fig. 7 B, F View Figure 7 ). Malar space granulate, 1.9 × as wide as base of mandible (Fig. 7 D, E View Figure 7 ). Occipital carina complete, joining hypostomal carina above base of mandible (Fig. 5 D View Figure 5 ); dorso-median part of occipital carina evenly arched (Fig. 7 C View Figure 7 ). Maxillary palp long, reaching epicnemial carina; fifth segment 4.0 × as long as mandible base. Dorsal length of eye 3.7 × as long as temple. Antenna with 36 antennomeres. Flagellum bristle-shaped, very long and slender (Fig. 7 A View Figure 7 ); apical flagellomere conical; first flagellomere 5.5 × as long as apical width and 1.4 × as long as second one.

Mesosoma. Mesosoma 1.6 × as long as high. Collar of pronotum long (Fig. 8 A, B View Figure 8 ). Pronotum indistinctly strigose on smooth surface laterally (Figs 7 D View Figure 7 , 8 A View Figure 8 ). Mesoscutum strongly convex (Fig. 8 A View Figure 8 ), with notaulus nearly absent, only impressed in the form of longitudinal strigose sculpture (Fig. 8 B View Figure 8 ); surface of mesoscutum granulate with scattered punctures except longitudinally strigose postero-median portion (Fig. 8 B View Figure 8 ). Epicnemial carina complete (Fig. 8 A View Figure 8 ); subtegular ridge sharp (Fig. 8 A View Figure 8 ); impression below speculum moderately deep (Fig. 8 A View Figure 8 ); sternaulus distinct, crenulate, sharp, almost reaching base of mid coxa (Fig. 8 A View Figure 8 ); posterior transverse carina of mesosternum absent; surface of mesopleuron obliquely strigose, smooth anteriorly, speculum slightly polished (Fig. 8 A View Figure 8 ). Scutellum highly elevated above metanotum, with crenulate lateral carina reaching apex and dorsal surface almost flat (Fig. 8 B View Figure 8 ); hind margin of metanotum with a triangular projection on each side of metanotum (Fig. 8 C, D View Figure 8 ). Juxtacoxal carina distinct, complete (Fig. 8 A View Figure 8 ). Propodeum in profile with area basalis flat and evenly slanted from base of area superomedia; area basalis not defined by carinae (Fig. 8 C View Figure 8 ); regular carinae of propodeum complete (Fig. 8 C, D View Figure 8 ); area superomedia hexagonal, 0.8 × as long as wide, posterior margin concave (Fig. 8 C View Figure 8 ); apex of area dentipara with strongly developed, moderately long and sharp apophysis, almost perpendicular to surface of propodeum, apophysis almost straight (Fig. 8 A View Figure 8 ); surface of propodeum mostly transversely strigose (Fig. 8 C View Figure 8 ); area petiolaris distinct laterally, transversely strigose (Fig. 8 C View Figure 8 ). Propodeal spiracle 1.5 × as long as wide.

Legs. Legs very slender and long (Fig. 7 A View Figure 7 ); all legs longer than fore wing. Hind femur and tibia 4.0, 7.0 × as long as maximum width respectively; ratio of length of tibiae fore: mid: hind = 1.0: 1.3: 2.7.

Wings. Fore wing 4.5 mm long; areolet 1.3 × as wide as long, 1.4 × as high as 3 rs-m, 0.7 × as high as 2 m-cu, pentagonal and sides narrowing anteriorly (Fig. 9 B View Figure 9 ); 1 cu-a opposite M & RS (Fig. 9 B View Figure 9 ); ramulus absent (Fig. 9 B View Figure 9 ). Hind wing with cu-a one-fourth as long as first abscissa of CU.

Metasoma. Metasoma mostly coriaceous (Figs 8 E, F View Figure 8 , 9 A View Figure 9 ). First tergite in profile almost straight, slightly bent downward at base of postpetiole (Fig. 8 E View Figure 8 ), smooth anteriorly, irregularly sculptured between latero-median carinae, coriaceous posteriorly; in dorsal view 4.5 × as long as wide at base (Fig. 8 F View Figure 8 ); petiole very long, rounded in cross-section, of approximately equal width and height (Fig. 8 E, F View Figure 8 ); in dorsal view petiole distinctly broadened posteriorly; latero-median carina present only on anterior portion of postpetiole (Fig. 8 F View Figure 8 ). Second tergite in dorsal view 1.8 × as long as wide anteriorly, distinctly rugose-punctate (Fig. 9 A View Figure 9 ). Gastrocoelus slightly impressed, in form of longitudinal groove (Fig. 9 A View Figure 9 ). Thyridium oval, distant from base of second tergite by 0.3 × its length (Fig. 9 A View Figure 9 ). Tergites 3–4 minutely punctate, 4–6 smooth with laterotergites separated by distinct creases (Fig. 9 A View Figure 9 ). Sternites 2–5 laterally sclerotized and middle part not sclerotized (Fig. 8 E View Figure 8 ). Hypopygium with apical margin rounded and longitudinally folded in middle. Ovipositor sheath with apex blunt, protruding beyond apex of metasoma (Fig. 8 E View Figure 8 ).

Colour. Body mostly yellow except for following: flagellomeres 2–6 (proximal half), 14–34, face except laterally, clypeus except basal half, frons anteriorly, ocellar area, vertex and occiput, temple, pronotum except laterally, propleuron, mesoscutum, metanotum, mesopleuron, wing venation, anterior half of second metasomal tergite, metasomal tergites 3–4, metasomal tergite 5 anteriorly dark brown to black, telotarsi, mid tarsus, hind femur and tibia distally, ovipositor sheath brown.

Male. Similar to female except: body length 6.9 mm, antenna with 35 antennomeres, face yellow except medio-longitudinally, clypeus completely yellow, frons medially black, mesopleuron with yellow patch posteriorly, area superomedia wider than long, second metasomal tergite completely yellow, flagellomeres 1–11, 15–33, brown, 12–14 yellow. Gonostyle narrow, ‘ finger-like’.

Variation.

One female paratype with wider longitudinal yellowish patch on face laterally and clypeus completely yellow.

Distribution.

Oriental region ( India).

Etymology.

APR dedicates this species in honor of Vedan (Hirandas Murali), a contemporary singer from Kerala whose work transcends musical boundaries and resonates across political and cultural divides. Vedan‘s music is noted for its capacity to inspire reflection, unity, and dialogue among individuals of differing political and cultural views, offering a sonic space where contrasting ideologies can intersect through shared emotional and aesthetic experience. This name pays tribute to the artist’s unique role in fostering connection and empathy in an increasingly polarized world.

Comparative diagnosis.

The new species is morphologically closer to H. brevispinus sp. nov., based on the shape of the area superomedia and colour of the frons. Apart from the differences cited in the identification key, H. vedani sp. nov. differs from H. brevispinus sp. nov. by the following: clypeus yellow in ventral half (dark brown in H. brevispinus sp. nov.), malar space 1.9 × as long as basal width of mandible (1.1 × in H. brevispinus sp. nov.) and first tergite 4.5 × as long as anterior width (5.1 × in H. brevispinus sp. nov.).