Guatteria confusa Maas & Westra

Maas, P. J. M., Westra, L. Y. T., Guerrero, S. Arias, Lobão, A. Q., Scharf, U., Zamora, N. A. & Erkens, R. H. J., 2015, Confronting a morphological nightmare: revision of the Neotropical genus Guatteria (Annonaceae), Blumea 60 (1), pp. 1-219 : 61

38. Guatteria confusa Maas & Westra — Map 10

Guatteria confusa Maas & Westra in Erkens et al. (2008) 481, f. 5. — Type: Díaz S. & Jaramillo 1478 (holo U; iso K, MO 2 sheets, WAG 2 sheets), Peru, Loreto, Prov. Maynas, Caserio de Urcumiraño, Río Napo , 120 m, 8 Oct. 1979.

Tree 4–16 m tall, diam not recorded; young twigs sparsely covered with appressed hairs, soon glabrous. Leaves: petiole 8–10 mm long, 2–3 mm diam; lamina narrowly elliptic, 17–24 by 5–9 cm (leaf index 2.9–3.4), coriaceous, not verruculose, shiny, blackish green above, pale brown below, glabrous above, sparsely covered with appressed hairs to glabrous below, base acute to obtuse, apex long-acuminate (acumen 10–35 mm long), primary vein impressed, flat or even slightly raised above, secondary veins distinct, 7–12 on either side of primary vein, raised above, smallest distance between loops and margin 3–5, tertiary veins raised above, reticulate. Flowers solitary in leaf axils; pedicels c. 3 mm long, c. 2 mm diam, fruiting pedicels 10–15 mm long, 2–4 mm diam, densely covered with ap- pressed, brown hairs, becoming glabrous in fruit, articulation at c. 0.3 from the base, bracts 4–5, soon falling, not seen, flower buds depressed ovoid; sepals free, broadly ovate-triangular, 5 –6 by 5– 6 mm, appressed, outer side densely covered with appressed, brown hairs; petals green to yellow in vivo (too young to be measured), outer side densely covered with ap- pressed, brownish white hairs; stamens not seen. Monocarps c. 20, green in vivo, black in sicco, ellipsoid, 19–23 by 8–9 mm, longitudinally wrinkled in sicco, glabrous, apex apiculate (apiculum 1–3 mm long), wall 0.5–0.7 mm thick, stipes 3–5 by 1–2 mm. Seed ellipsoid, 15–20 by 7–8 mm, dark brown, apex distinctly apiculate (apiculum 1–2 mm long), longitudinally and transversely grooved, raphe not distinct from rest of seed.

Distribution — Amazonian Peru (Loreto).

Habitat & Ecology — In non-inundated forest, on clayey soil. At elevations of 120– 170 m. Flowering: June, July, November; fruiting: June to August, October.

Vernacular names — Anona (Rimachi Y. 1059), Espintana (Rimachi Y. 2458, 4033).

Note — The leaves of G. confusa are somewhat suggestive of Ephedranthus because of a comparatively low number of secondary veins.