Gamblea malayana (M.R. Henderson) C .- B . Shang, Lowry & Frodin, 2000

Shang, C. - B., Lowry Ii, Porter P. & Frodin, David G., 2000, A taxonomic revision and re-definition of the genus Gamblea (Araliaceae), Adansonia (3) 22 (1), pp. 45-55 : 53

3. Gamblea malayana (M.R. Henderson) C.- B. Shang, Lowry & Frodin , comb. nov.

Acanthopanax malayanus M.R. Henderson , “ malayana ”, Gard. Bull. Straits Settlem. 7: 105, pl. 23 (1933); W.R. Philipson, Fl. Malesiana, ser. 1, 9: 103 (1979). — Eleutherococcus malayanus (M.R. Henderson) B.C. Stone, Malaysian For. 43: 395 (1980). — Evodiopanax malayanus (M.R. Henderson) C.- B. Shang & J.- Y. Huang, J. Nanjing For. Univ. 17: 34 (1993).

— Type: M.R. Henderson SF 23476, Malaysia, Pahang, Cameron’s Highlands, beyond Tanah Rata , 1430 m, 7 Apr. 1930 (holo-, SING; iso-, K [6 sheets]!, NY!).

Trees 10-15(-25) m tall, 15-100 cm dbh. Leaves 3-5-foliolate, central leaflet narrowly elliptic to slightly ovate, (6.5-)9-17 × (2-) 2-5.5 cm at maturity, 3-4.5 times as long as wide, with 8-10 secondary veins on each side of the midvein, domatia occasionally absent or more often obscure on abaxial surface at base of secondary veins, ca. 0.2 mm in diam., tertiary veins distinctly raised on abaxial surface, margins subentire, with evident, divergent, ciliate-hispid teeth (1-) 1.5-2.5 mm long, spaced every 1-2 mm. Inflorescence a compound umbel, primary axes (2-)3-7, unbranched, 4.5-11 cm long at anthesis and in fruit, each terminating in an umbellule (rarely also with a single lateral umbellule); calyx a narrow rim, with 4-5(-6) small, triangular teeth; styles 2, united for 2/3-3/4 of their length. Fruit globose to obloid, 6-7 × 7-10 mm, usually somewhat compressed laterally, disk 1.5-2 mm in diam., with a narrow, entire rim and usually 4-5 (-6) persistent calyx teeth.

Gamblea malayana occurs in moist montane forest between about 1400 and 2500 m elevation in the Cameron Highlands of peninsular Malaysia and in northern Sumatra. It is easily distinguished from other species of Gamblea by its narrowly elliptic leaflets with subentire margins bearing well developed ciliate-hispid teeth usually 1.5-2.5 mm long, and its laterally compressed,globose to obloid fruit measuring 6-7 × 7-10 mm.

MATERIAL EXAMINED. — INDONESIA: Sumatra, de Wilde & de Wilde-Duyfjes 16725, Atjeh, Gunung Leuser Nature Reserve, Gunung Mamas, ca. 16 km SW from mouth of Lau Ketambe, ca. 30 km NW of Kutatjane, 1900 m, 8 May 1975, fl. ( K), 16856, same locality, ca. 23 km SW from mouth of Lau Ketambe, 2500-2600 m, 12 May 1975, fr. ( K, MO). MALAYSIA: Pahang, Carrier SF 27650, Birchang, Cameron Highlands, 9 Aug. 1933, fr. ( K); W.-L. Chew 829, Bukit Ruil., Cameron Highlands, 4°30’N, 101°21’E, 1700 m, 6 Oct. 1963, fr. ( K); M R. Henderson SF 23476, Cameron’s Highlands, beyond Tanah Rata, 1430 m, 7 Apr. 1930 ( K [6 sheets], NY, SING); Holttum SF 31368, Sungai Burong, Cameron Highlands, 1500 m, 20 May 1936, y.fr. ( K [3 sheets]); Kochummen KEP FRI 19059, path to G. Jasar, Cameron Highlands, 1600 m, 23 Aug. 1977, ster. ( K); K. Ogata KEP 110312, G. Jasar, Cameron Highlands, 1500 m, 2 Mar. 1968, fr. ( K); Stone 14466, G. Beremban, near summit, Cameron Highlands, 1700 m, 7 Mar. 1980, bud ( K); Symington SF 36080, edge of Tama Sedia, Cameron Highlands, 6 Apr. 1934, fl. ( K [2 sheets]), SF 36216, golf course, Cameron Highlands, 11 Apr. 1934, fl. ( K [2 sheets]); K.M. Wong KEP FRI 35250, Gunung Jasar, Cameron Highlands, 1500 m, 13 Aug. 1986, fr. ( K).