Gamblea ciliata C.B. Clarke var. ciliata

Shang, C. - B., Lowry Ii, Porter P. & Frodin, David G., 2000, A taxonomic revision and re-definition of the genus Gamblea (Araliaceae), Adansonia (3) 22 (1), pp. 45-55 : 49

1a. Gamblea ciliata C.B. Clarke var. ciliata

Acanthopanax evodiaefolius Franch. var. ferrugineus W.W. Smith , Notes Bot. Gard. Edinburgh 10: 6 (1917). — Evodiopanax evodiaefolius (Franch.) Nakai var. ferrugineus (W.W. Smith) Nakai , “ ferrugineum ”, J. Arnold Arbor. 5: 8 (1924). — Evodiopanax ferrugineus (W.W. Smith) Grushv. & Skvorts., Novosti Sist. Vyssh. Rast. 22: 177 (1985).

— Lectotype (here designated): Forrest 12068, China, Yunnan, Shweli–Salwin divide, 25°20’N, Aug. 1913, fl. ( E!; iso-, E!, K!); syn. nov.

Acanthopanax evodiaefolius Franch. var. gracilis W.W. Smith , Notes Bot. Gard. Edinburgh 10: 6 (1917). — Evodiopanax gracilis (W.W. Smith) Grushv. & Skvorts., Novosti Sist. Vyssh. Rast. 22: 177 (1985).

— Type: Forrest 11282, China, Yunnan, Lichiang Range , 27°40’N, Sep. 1913, fr. (holo-, E!; iso-, E!, K!); syn. nov.

Acanthopanax evodiaefolius Franch. var. glaucus Feng, Fl. Yunnanica 2: 486 (1979).

— Type: T. T. Yu 19318, China, Yunnan, Salwin–Kiukian divide, Newahlung , 3500 m, 11 July 1938, fl. (holo-, KUN!; iso-, E!, K!); syn. nov.

Flowers and fruits with (2-)3-4(-5) style branches, largest leaflets (8-) 10-20 cm long, with (6-)8-14 secondary veins on each side of the midvein, fruit with a disk (1-) 1.5-3 mm in diam.

SMITH (1917) cited two collections when he described Acanthopanax evodiaefolius var. ferrugineus , which therefore represent syntypes. One of these (Forrest 12068) comprises material in full flower, and we have selected the lectotype from among the two specimens of this gathering on deposit in Edinburgh.

MATERIAL EXAMINED. — NEPAL: Beer 25556, Iswa Khola, 3350 m, 5 Oct. 1975, ster. (BM); Bowes Lyon 173, Ankhu Khola, Barang, 2600 m, 5 Apr. 1962, bud (BM), 2012, Lumding Khola, Dudh Kosi, 3000 m, 8 June 1964, bud (BM); Dobremez DBR.NEP 1398, Tapletok, 27°27’N, 87°50’E, 3300 m, 6 Oct. 1971, fr. (BM); Noshiro et al. 9263208, Mechi zone, Taplejung Dist., Chairam-Andha Phedi-Dorongen, 27°31’38’’- 27°35’02’’N, 87°54’46’’- 87°58’11’’E, 2890-3720 m, 11 June 1992, bud (BM); Ohba et al. 54098, Koshi zone, Sankhuwa Sabha Dist., Milke Danda, Angare Kharka-Chhippon, 27°20’N, 87°30’E, 2930-2980 m, 18 July 1991, y.fr. (BM); Polunin 430, Langtang, 3200 m, June 1949, fl. (BM, E); Stainton 661 Arum Valley, Barum Khola, N of Num, 2730 m (BM, E); M. Suzuki et al. 8860365, Koshi zone, Sankhuwa Sabha Dist., Phemathang Kharka-Barun Khola-Numbuk Cha Ding Kharka, 27°40’N, 87°10’E, 3300 m, 29 July 1988, y.fr. (BM), 8880451, Koshi zone, Sankhuwa Sabha Dist., Tashi Gaun, Uttise Kharka-Bhainsi Kharka, 27°35’N, 87°15’E, 2160-2540 m, 14 July 1988, y.fr. (BM); H. J. Williams 627, Yiringdham, 27°20’N, 87°57’E, 3230 m, 16 June 1969, fl. (BM), 966, Yamphodin, 27°27’N, 87°57’E, 3050 m, 25 June 1969, fr. (BM). INDIA: Assam, Kingdon Ward 8171, Delei valley, 28°20’ N, 96°37’E, 2150-2750 m, 6 May 1928, fl. ( K). Manipur, Watt 6889, Japvo, 3150 m, 15 May 1882, ster. ( K). Sikkim, Cave s.n., Tonglu, 2750 m, 15 June 1912, fl. ( E), Chowhanjan, 3350 m, 14 Aug. 1925, y.fr. ( E [2 sheets]); C. B. Clarke 25864, 26034 A, Jongri, 15 Oct. 1875, ster. ( K), 26034 B, Jongri, 3650 m, ster. (BM), 41359, Jakpho, 3020 m, 25 Oct. 1885, ster. ( K); Gamble 25 A, Tongloo, 2750 m, 5 Oct. 1875, fr. ( K), 25 B, Tonglo, 3050 m, 5 Oct. 1875, fr. ( K), 3039, Tonglo, 3050 m, Nov. 1874, ster. ( K), 10412, Tonglo, 3050 m, July 1882, fl. (BM, K); Gammie 614, Tankra Mt., 3050 m, 5 Aug. 1892, y.fr. (P); J.D. Hooker s.n., without precise locality, 3050 m, bud ( K); G. King 187, without precise locality, fr. (BM), without precise locality, 1874-5, ster. (P); O. Kuntze 6954, without precise locality, 2450 m, Nov. 1875, fr. (NY); Kurz s.n., without precise locality, 14 Oct. 1868, fr. (BM); Lace s.n., Tonglu, 3050 m, June 1902, bud ( E); Leonige 8563, Tonglo, 3050 m, Oct. 1880, fr. ( K); Ribu & Rhomoo 6344, Chowbhanjan, 3350 m, 1913, fr. ( E); Rohmoo Lepeha 1230, Chowbhanjan, 3050 m, 12 Aug. 1913, fl. ( E). West Bengal, Haines 1108, Pankasari, 2450 m, 1904, fr. ( K). BHUTAN: Cooper 2745, Parshong Lumpu, 3050 m, 28 July 1914, y.fr. (BM, E), 2962, Dotena Limpu, 2750 m, 1 Oct. 1914, fr. (BM, E), 3992, Kopub Pumthang, 2750 m, 18 June 1995, fl. (BM, E), 4536, Tarkigong, 2750 m, 23 Aug. 1915, fr. ( E), 4636, Lashigong Kurmed, 2750 m, 23 Aug. 1915, fr. (BM); Frei-Pont 3, Bhumthang Dist., Lami Gompa, 27°33’N, 90°42’E, 30 Aug. 1984, fr. ( E); Grierson & Long 1899, above Sengor, NW of Mongar, 27°23’N, 91°01’E, 3150 m, 14 June 1979, fr. ( E, K); Ludlow et al. 20487, Shingbe (Me La), 3350 m, 5 July 1949, fl. (BM); Sargent 200, Tongsa Dist., Singkma, 27°30’N, 90°33’E, 13 Aug. 1983, ster. ( E). MYANMAR: Kingdon-Ward 1707, Naung Chaung, Nwai Dwide, 2750-3050 m, 24 June 1914, fl. ( E), 12952, hills E of the Nam Tawai, 27°50’N, 97°50’E, 2750-2900 m, 18 Aug. 1937, fr. (BM), 13057, without precise locality, fr. (BM), 20859, North Triangle, Wing Bum, above Ahkail, 2900-3050 m, 14 May 1953, fl. (BM, E), 21093, same locality, 1 July 1953, fl. (BM); Cooper et al. 3992, without precise locality (BM, E), 4536 (BM). CHINA: Xizang, Chen Shu-zhi et al. 385, Me Tuo to Bo Mi, 22 Aug. 1982 (PE), Chen Shu-zhi 739, Me Tuo (NFU); Forrest 14940, Doku-la, Mekong-Salwin divide, Sep. 1917, fr. ( E); Kingdon Ward 19641, Tha Chu valley, 2750 m, 15 July 1950, fl. (BM, E); Li Pu-sheng 385, Me Tuo, 2800 m (NFU), 6726, Bo Mi, 2800 m, (PE), 7164, Cha Yu xian, 2600-3100 m, (NFU); Qing Zhan Exped. 6982, Bo Mi, 3150 m, 19 July 1975 (PE), 6986, Jilong, 3150 m, 19 July 1975 (PE), 73613, Cha Yu xian, 3000 m, 1 Aug. 1974 (PE), 751290, Lin Zhi, 3200 m, 4 Aug. 1975 (PE); Qing Zhan Medicin. Exped. 282, Jilong, 3300 m, 16 June 1972 (PE); Rock 22283, Solo-la, 2750 m, May-June 1932, fl. ( E, K), 22652, same locality, Aug.-Oct. 1932, fr. ( E); Shu Feng-qiang 7813, Lou Lang, 3100-3400 m (NFU); G.G.Tang 1447, 1599, 10001, Lou Lang, 3300 m (NFU); Ying Jun-sheng et al. 65123, Bo Mi, 27 Aug. 1969 (PE); Zhang Yong-lian et al. 3532, Nie La Mu, 18 May 1966 (PE). Sichuan, Farges 700, Dist. Tchen-keou-tin, 1400 m, July, fl., fr. (P [3 sheets]), 1400, same locality, fl. ( K), Farges s.n., same locality, fl. (P [3 sheets], TCD); Rock 24394, Siga Shan, 3350-3650 m, 1932, fr. ( E, K). Yunnan, Bartholomew et al. 1147, Dali xian, fr. ( E); Delavay s.n., same locality, 7 Aug. 1888, fr. (P); K. M. Feng 7586, Gon Shan, 7 Sep. 1940 (PE); Forrest 8887, Shweli-Salwin divide, 25°20’N, 2150 m, Aug. 1912, fr. ( E, K), 11282, same locality, Sep. 1913, fr. ( E [2 sheets], K), 12068, same locality, Aug. 1913, fl. ( E [2 sheets], K), 15922, same locality, 2150 m, Sep. 1917, fl. ( E, K, P), 19722, Mekong-Salwin divide, 27°30’N, 98°56’E, 2750 m, July 1921, fl. ( E, K, P), 29037, without precise locality, fr. ( E); Handel-Mazzetti 8329, Doyonlumba valley, Lu-djiang Riven (Salween), 2500- 3450 m, 23 Sep. 1915, fr. ( K); Kingdon Ward 790, Mekong-Salween divide, 3350 m, 17 July 1913, fl. ( E); Qing Zhan Exped. 7501, Gon Shan, 25 June 1982 (PE); Rock 4133, Yangtze watershed, W slopes of Likiang Snow Range, 3505- 3350 m, 6 June 1922, fl. ( E), 5106, between Likiang, Youngning and Youngpei, route to Mili, May-June 1922, fl. ( A), 9512, Lotue Shan, Labako mountains, W of Yangtze bend at Shiku, June 1923, fl. ( E); Schneider 2471, Lichiang, 3300 m, 9 Sep. 1914, fr. ( K), 3258, near Li Chiang, 3000 m, 6 Oct. 1914, fr. ( K); Sino-Amer. Bot. Exped. 292, Yangbi Xian, W side of Diancang Shan range, vicinity of Dapingzi, 25°43’N, 100°02’E, 3000 m, 19 June 1984, y.fr. ( A, E), 651, same locality, 25°50’N, 99°59’E, 2600-3100 m, 1 July 1984, y.fr. ( A, E), 1147, Dali Xian, Diancang Shan range, 25°53’N, 100°01’E, 2900-3300 m, 18 July 1984, fr. ( A, E); Su Sou-gui 4685, Jin Tong Xian, 2600 m, April 1959 (PE); S. Ten 508, Kau ty, near Pe yen tsin, 19 May 1918, fl. ( E), S. Ten s.n., same locality, 15 Apr. 1916, fl. (P), 19 May 1917, fl. (P); H. T. Tsai 51246, Liang Shan, La’mi, 2200 m, 9 Aug. 1932, fr. ( A); T. T. Yu 7852, Antuntze, Dokerla, 3100 m, 4 Nov. 1937, fr. ( E), 8426, Atuntze, Mt. Kaakerpu, 3200 m, 3 July 1937, fl. ( E), 10488, same locality, 3000 m, 27 Sep. 1937, fr. ( E), 10558, Antuntze, Mt. Miyetzimu, 3200 m, 16 Oct. 1937, fr. ( E), 11766, without precise locality, fl. ( A), 15980, Shunning, 2900 m, 26 May 1938, fl. ( A); 19318, Salwin-Kiukian divide, Newahlung, 3500 m, 11 July 1938, fl. ( E, K, KUN), 19445, Kiukiang Valley (Taron), Chiengen, 1700 m, 26 July 1938, fr. ( E), 20258, Salwin-Kiukiang divide, Lunguailakam 3300 m, 14 Sep. 1938, fr. ( E).