Gamblea ciliata C.B. Clarke var. ciliata

Shang, C. - B., Lowry Ii, Porter P. & Frodin, David G., 2000, A taxonomic revision and re-definition of the genus Gamblea (Araliaceae), Adansonia (3) 22 (1), pp. 45-55 : 49

1a. Gamblea ciliata C.B. Clarke var. ciliata

Acanthopanax evodiaefolius Franch. var. ferrugineus W.W. Smith , Notes Bot. Gard. Edinburgh 10: 6 (1917). — Evodiopanax evodiaefolius (Franch.) Nakai var. ferrugineus (W.W. Smith) Nakai , “ ferrugineum ”, J. Arnold Arbor. 5: 8 (1924). — Evodiopanax ferrugineus (W.W. Smith) Grushv. & Skvorts., Novosti Sist. Vyssh. Rast. 22: 177 (1985).

— Lectotype (here designated): Forrest 12068, China, Yunnan, Shweli–Salwin divide, 25°20’N, Aug. 1913, fl. ( E!; iso-, E!, K!); syn. nov.

Acanthopanax evodiaefolius Franch. var. gracilis W.W. Smith , Notes Bot. Gard. Edinburgh 10: 6 (1917). — Evodiopanax gracilis (W.W. Smith) Grushv. & Skvorts., Novosti Sist. Vyssh. Rast. 22: 177 (1985).

— Type: Forrest 11282, China, Yunnan, Lichiang Range , 27°40’N, Sep. 1913, fr. ( holo-, E!; iso-, E!, K!); syn. nov.

Acanthopanax evodiaefolius Franch. var. glaucus Feng, Fl. Yunnanica 2: 486 (1979).

— Type: T. T. Yu 19318, China, Yunnan, Salwin–Kiukian divide, Newahlung , 3500 m, 11 July 1938, fl. ( holo-, KUN!; iso-, E!, K!); syn. nov.

Flowers and fruits with (2-)3-4(-5) style branches, largest leaflets (8-) 10-20 cm long, with (6-)8-14 secondary veins on each side of the midvein, fruit with a disk (1-) 1.5-3 mm in diam.

SMITH (1917) cited two collections when he described Acanthopanax evodiaefolius var. ferrugineus , which therefore represent syntypes. One of these ( Forrest 12068) comprises material in full flower, and we have selected the lectotype from among the two specimens of this gathering on deposit in Edinburgh.

MATERIAL EXAMINED. — NEPAL: Beer 25556, Iswa Khola, 3350 m, 5 Oct. 1975, ster. (BM); Bowes Lyon 173, Ankhu Khola, Barang, 2600 m, 5 Apr. 1962, bud (BM), 2012, Lumding Khola, Dudh Kosi, 3000 m, 8 June 1964, bud (BM); Dobremez DBR.NEP 1398, Tapletok, 27°27’N, 87°50’E, 3300 m, 6 Oct. 1971, fr. (BM); Noshiro et al. 9263208, Mechi zone, Taplejung Dist., Chairam-Andha Phedi-Dorongen, 27°31’38’’- 27°35’02’’N, 87°54’46’’- 87°58’11’’E, 2890-3720 m, 11 June 1992, bud (BM); Ohba et al. 54098, Koshi zone, Sankhuwa Sabha Dist., Milke Danda, Angare Kharka-Chhippon, 27°20’N, 87°30’E, 2930-2980 m, 18 July 1991, y.fr. (BM); Polunin 430, Langtang, 3200 m, June 1949, fl. (BM, E); Stainton 661 Arum Valley, Barum Khola, N of Num, 2730 m (BM, E); M. Suzuki et al. 8860365, Koshi zone, Sankhuwa Sabha Dist., Phemathang Kharka-Barun Khola-Numbuk Cha Ding Kharka, 27°40’N, 87°10’E, 3300 m, 29 July 1988, y.fr. (BM), 8880451, Koshi zone, Sankhuwa Sabha Dist., Tashi Gaun, Uttise Kharka-Bhainsi Kharka, 27°35’N, 87°15’E, 2160-2540 m, 14 July 1988, y.fr. (BM); H. J. Williams 627, Yiringdham, 27°20’N, 87°57’E, 3230 m, 16 June 1969, fl. (BM), 966, Yamphodin, 27°27’N, 87°57’E, 3050 m, 25 June 1969, fr. (BM). INDIA: Assam, Kingdon Ward 8171, Delei valley, 28°20’ N, 96°37’E, 2150-2750 m, 6 May 1928, fl. ( K). Manipur, Watt 6889, Japvo, 3150 m, 15 May 1882, ster. ( K). Sikkim, Cave s.n., Tonglu, 2750 m, 15 June 1912, fl. ( E), Chowhanjan, 3350 m, 14 Aug. 1925, y.fr. ( E [2 sheets]); C. B. Clarke 25864, 26034 A, Jongri, 15 Oct. 1875, ster. ( K), 26034 B, Jongri, 3650 m, ster. (BM), 41359, Jakpho, 3020 m, 25 Oct. 1885, ster. ( K); Gamble 25 A, Tongloo, 2750 m, 5 Oct. 1875, fr. ( K), 25 B, Tonglo, 3050 m, 5 Oct. 1875, fr. ( K), 3039, Tonglo, 3050 m, Nov. 1874, ster. ( K), 10412, Tonglo, 3050 m, July 1882, fl. (BM, K); Gammie 614, Tankra Mt., 3050 m, 5 Aug. 1892, y.fr. (P); J.D. Hooker s.n., without precise locality, 3050 m, bud ( K); G. King 187, without precise locality, fr. (BM), without precise locality, 1874-5, ster. (P); O. Kuntze 6954, without precise locality, 2450 m, Nov. 1875, fr. (NY); Kurz s.n., without precise locality, 14 Oct. 1868, fr. (BM); Lace s.n., Tonglu, 3050 m, June 1902, bud ( E); Leonige 8563, Tonglo, 3050 m, Oct. 1880, fr. ( K); Ribu & Rhomoo 6344, Chowbhanjan, 3350 m, 1913, fr. ( E); Rohmoo Lepeha 1230, Chowbhanjan, 3050 m, 12 Aug. 1913, fl. ( E). West Bengal, Haines 1108, Pankasari, 2450 m, 1904, fr. ( K). BHUTAN: Cooper 2745, Parshong Lumpu, 3050 m, 28 July 1914, y.fr. (BM, E), 2962, Dotena Limpu, 2750 m, 1 Oct. 1914, fr. (BM, E), 3992, Kopub Pumthang, 2750 m, 18 June 1995, fl. (BM, E), 4536, Tarkigong, 2750 m, 23 Aug. 1915, fr. ( E), 4636, Lashigong Kurmed, 2750 m, 23 Aug. 1915, fr. (BM); Frei-Pont 3, Bhumthang Dist., Lami Gompa, 27°33’N, 90°42’E, 30 Aug. 1984, fr. ( E); Grierson & Long 1899, above Sengor, NW of Mongar, 27°23’N, 91°01’E, 3150 m, 14 June 1979, fr. ( E, K); Ludlow et al. 20487, Shingbe (Me La), 3350 m, 5 July 1949, fl. (BM); Sargent 200, Tongsa Dist., Singkma, 27°30’N, 90°33’E, 13 Aug. 1983, ster. ( E). MYANMAR: Kingdon-Ward 1707, Naung Chaung, Nwai Dwide, 2750-3050 m, 24 June 1914, fl. ( E), 12952, hills E of the Nam Tawai, 27°50’N, 97°50’E, 2750-2900 m, 18 Aug. 1937, fr. (BM), 13057, without precise locality, fr. (BM), 20859, North Triangle, Wing Bum, above Ahkail, 2900-3050 m, 14 May 1953, fl. (BM, E), 21093, same locality, 1 July 1953, fl. (BM); Cooper et al. 3992, without precise locality (BM, E), 4536 (BM). CHINA: Xizang, Chen Shu-zhi et al. 385, Me Tuo to Bo Mi, 22 Aug. 1982 (PE), Chen Shu-zhi 739, Me Tuo (NFU); Forrest 14940, Doku-la, Mekong-Salwin divide, Sep. 1917, fr. ( E); Kingdon Ward 19641, Tha Chu valley, 2750 m, 15 July 1950, fl. (BM, E); Li Pu-sheng 385, Me Tuo, 2800 m (NFU), 6726, Bo Mi, 2800 m, (PE), 7164, Cha Yu xian, 2600-3100 m, (NFU); Qing Zhan Exped. 6982, Bo Mi, 3150 m, 19 July 1975 (PE), 6986, Jilong, 3150 m, 19 July 1975 (PE), 73613, Cha Yu xian, 3000 m, 1 Aug. 1974 (PE), 751290, Lin Zhi, 3200 m, 4 Aug. 1975 (PE); Qing Zhan Medicin. Exped. 282, Jilong, 3300 m, 16 June 1972 (PE); Rock 22283, Solo-la, 2750 m, May-June 1932, fl. ( E, K), 22652, same locality, Aug.-Oct. 1932, fr. ( E); Shu Feng-qiang 7813, Lou Lang, 3100-3400 m (NFU); G.G.Tang 1447, 1599, 10001, Lou Lang, 3300 m (NFU); Ying Jun-sheng et al. 65123, Bo Mi, 27 Aug. 1969 (PE); Zhang Yong-lian et al. 3532, Nie La Mu, 18 May 1966 (PE). Sichuan, Farges 700, Dist. Tchen-keou-tin, 1400 m, July, fl., fr. (P [3 sheets]), 1400, same locality, fl. ( K), Farges s.n., same locality, fl. (P [3 sheets], TCD); Rock 24394, Siga Shan, 3350-3650 m, 1932, fr. ( E, K). Yunnan, Bartholomew et al. 1147, Dali xian, fr. ( E); Delavay s.n., same locality, 7 Aug. 1888, fr. (P); K. M. Feng 7586, Gon Shan, 7 Sep. 1940 (PE); Forrest 8887, Shweli-Salwin divide, 25°20’N, 2150 m, Aug. 1912, fr. ( E, K), 11282, same locality, Sep. 1913, fr. ( E [2 sheets], K), 12068, same locality, Aug. 1913, fl. ( E [2 sheets], K), 15922, same locality, 2150 m, Sep. 1917, fl. ( E, K, P), 19722, Mekong-Salwin divide, 27°30’N, 98°56’E, 2750 m, July 1921, fl. ( E, K, P), 29037, without precise locality, fr. ( E); Handel-Mazzetti 8329, Doyonlumba valley, Lu-djiang Riven (Salween), 2500- 3450 m, 23 Sep. 1915, fr. ( K); Kingdon Ward 790, Mekong-Salween divide, 3350 m, 17 July 1913, fl. ( E); Qing Zhan Exped. 7501, Gon Shan, 25 June 1982 (PE); Rock 4133, Yangtze watershed, W slopes of Likiang Snow Range, 3505- 3350 m, 6 June 1922, fl. ( E), 5106, between Likiang, Youngning and Youngpei, route to Mili, May-June 1922, fl. ( A), 9512, Lotue Shan, Labako mountains, W of Yangtze bend at Shiku, June 1923, fl. ( E); Schneider 2471, Lichiang, 3300 m, 9 Sep. 1914, fr. ( K), 3258, near Li Chiang, 3000 m, 6 Oct. 1914, fr. ( K); Sino-Amer. Bot. Exped. 292, Yangbi Xian, W side of Diancang Shan range, vicinity of Dapingzi, 25°43’N, 100°02’E, 3000 m, 19 June 1984, y.fr. ( A, E), 651, same locality, 25°50’N, 99°59’E, 2600-3100 m, 1 July 1984, y.fr. ( A, E), 1147, Dali Xian, Diancang Shan range, 25°53’N, 100°01’E, 2900-3300 m, 18 July 1984, fr. ( A, E); Su Sou-gui 4685, Jin Tong Xian, 2600 m, April 1959 (PE); S. Ten 508, Kau ty, near Pe yen tsin, 19 May 1918, fl. ( E), S. Ten s.n., same locality, 15 Apr. 1916, fl. (P), 19 May 1917, fl. (P); H. T. Tsai 51246, Liang Shan, La’mi, 2200 m, 9 Aug. 1932, fr. ( A); T. T. Yu 7852, Antuntze, Dokerla, 3100 m, 4 Nov. 1937, fr. ( E), 8426, Atuntze, Mt. Kaakerpu, 3200 m, 3 July 1937, fl. ( E), 10488, same locality, 3000 m, 27 Sep. 1937, fr. ( E), 10558, Antuntze, Mt. Miyetzimu, 3200 m, 16 Oct. 1937, fr. ( E), 11766, without precise locality, fl. ( A), 15980, Shunning, 2900 m, 26 May 1938, fl. ( A); 19318, Salwin-Kiukian divide, Newahlung, 3500 m, 11 July 1938, fl. ( E, K, KUN), 19445, Kiukiang Valley (Taron), Chiengen, 1700 m, 26 July 1938, fr. ( E), 20258, Salwin-Kiukiang divide, Lunguailakam 3300 m, 14 Sep. 1938, fr. ( E).