Cosmopolitodus hastalis ( Agassiz, 1838 )

Szabó, Márton, 2019, Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum, Fragmenta Palaeontologica Hungarica 36, pp. 53-90 : 65

Cosmopolitodus hastalis ( Agassiz, 1838)

( Figs 66–71 )

Referred material – 13 teeth (V.61.917, V.61.925, V.61.1278, V.61.1280, V.62.395, V.62.397, V.62.399, V.69.286, V.69.306, V.69.1010, V.69.1038, V.69.1092).

Description – The crown of upper anterior and anterolateral teeth ( Figs 66, 67 ) is high, triangular and labiolingually flattened. The labial crown face is flat, while the lingual is weakly convex. Upper lateral teeth have a distally directed crown, rapidly widening basally, extending onto the shoulders of the root lobes, and sometimes forming a heel in upper distal files (see V.62.399, Figs 68, 69 ). The short root lobes of upper files are weakly pronounced lingual protuberance. Lower teeth ( Figs 70, 71 ) have lingually weakly curved, narrow crown (compared to upper teeth), the labial crown face is weakly, while the lingual is strongly convex. The root of lower files is built up of well-separating lobes. The crown faces and cutting edges of all files are smooth. The lingual crown-root boundary is marked by a narrow dental band in all files.

Remarks – This species is widespread in the Badenian deposits of the Central Paratethys ( SZABÓ & KOCSIS 2016, table 2). Following the dental similarities of Cosmopolitodus and Carcharodon carcharias , some authors place the species into the genus Carcharodon (e.g. EHRET et al. 2012; TRIF & CODREA 2017; KENT in GODFREY 2018; MAISCH et al. 2018). Since the evolutional relationship of Cosmopolitodus and Carcharodon is not fully resolved at present, here I follow CAPPETTA (2012), who places species hastalis in genus Cosmopolitodus .

Figures

Figs 66–71. Cosmopolitodus hastalis (Agassiz, 1838). – Fig. 66. Upper anterior tooth, V.69.1010, lingual view. – Fig. 67. Upper anterolateral tooth, V.61.1280, lingual view. – Figs 68–69. Upper lateral tooth, V.62.399. – Fig. 68. Labial view. – Fig. 69. Close-up of the mesial crown edge of the same specimen. – Fig. 70. Lower anterior tooth, V.69.1038, lingual view. – Fig. 71. Lower anterior tooth, V.69.306, lingual view. – Fig. 72. Isurus oxyrinchus (Agassiz, 1843),?lower anterior tooth, V.61.915, labial view. Scale bars: 10 mm