Clivina opacielytra, Balkenohl, 2025

Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), Zootaxa 5592 (1), pp. 1-70 : 24-25

publication ID	https://doi.org/10.11646/zootaxa.5592.1.1
publication LSID	lsid:zoobank.org:pub:0E478171-152D-4A6F-9CB5-D9C9B5ABB744
DOI	https://doi.org/10.5281/zenodo.14953716
persistent identifier	https://treatment.plazi.org/id/03F487F9-7302-FFB7-53D4-A19FFF1BF9B9
treatment provided by	Plazi
scientific name	Clivina opacielytra
status	sp. nov.

Treatment

Clivina opacielytra sp. nov.

( Figs 13 View FIGURES 3–14 , 48 View FIGURES 38–52 , 111 View FIGURES 110–128 )

Type material. Holotype: ♀, with labels and data: white, black printed “ INDONESIEN Samabusa 25 km östl. Nabire COROVA - RIVER leg. Dr. D. DASCHE 14.-18.02.95“ / “ Clivina group helferi det. Balkenohl, 1999 ”, “COLL. WRASE BERLIN” / green, black printed “COLL. WRASE GUSOW-PLATKOW GERM.” ( CDWGP) . Paratype: 1 ♀, with labels and data: all white, black printed “Palmer R. at Black R. Papua July 22-31, 1936 ArchboldExped.” / “Borrowed fr AMNH 1961” / “ Clivina wallacei Pz. det. Darl. 68” / barcode “MCZ-ENT 00806446” ( MCZ) .

Remarks. The paratype shows several damages: the two right terminal antennomeres are missing. In both of the front legs, the tarsomeres are missing. The right intermediate leg is missing as well as three tarsomeres of the left hind leg, and the left tibia with tarsomeres is dislocated. These damages gave reason to select the complete specimen as holotype.

The paratype designated is mentioned in Darlington (1971) under the name Clivina wallacei Putzeys as the “first surely recorded” C. wallacei from the island. The whereabouts of the other two specimens mentioned in Darlington (1971) remain unclear but should be investigated in order to clear occurrence of C. wallacei wallacei in Papua/New Guinea.

Description. Measurements. Holotype: Body length 5.47 mm, width 1.52, ratio L/W of pronotum 0.99, ratio L/W of elytra 1.95. Paratype: Body length 5.31 mm, width 1.46, ratio L/W of pronotum 0.99, ratio L/W of elytra 2.0. Colour fulvous. Head and pronotum glossy, elytra opaque. Lateral margins of head fuscous. Antennae, mouthparts, intermediate and hind legs yellowish.

Head about a fifth narrower than pronotum. Clypeus with central part straight, projection of lateral teeth distinct, separated from clypeal-wing by distinct notch; wing distinct, less high as lateral teeth but as high as central part of clypeus, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin regularly convex, clypeus, wing and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a blunt keel, separated from wing by narrow furrow. Clypeus transverse, convex, smooth, clypeal-furrow wide, flat, with indistinct reticulation, frons moderately convex, with scattered small punctures, with pore at middle, separated from supraantennal plate and supraorbital carina by moderately wide furrow, with clypeal seta in front; supraorbital carinae narrow, with two supraorbital setigerous punctures at middle of eye and at posterior gena-level. Neck constriction closed, developed as step from frons, with punctures posteriorly. Eye prominent, gena distinct, convex posteriorly. Antenna long, reaching over base of elytron, antennomeres four to ten elongate (L/ W 2.0). Labrum pentagonal, anteriorly with flat angle and pointed at middle, irregularly to isodiametrically reticulated, six-setose. Lobe of mentum flat, regularly oval at tip, median tooth about as high as lobe, bent dorsally, rounded anteriorly.

Pronotum ( Fig. 48 View FIGURES 38–52 ) with disk flat in lateral view, in frontal view moderately convex laterally, flattened at middle. Shape peltate, seemingly longer than wide but measured as long as wide. Reflexed lateral margin smooth, slightly convex, widest at middle; anterior angle rounded but distinct, posterior one widely rounded, with tooth indistinct; lateral channel narrow, reticulated; margin continuing up to base as sharp, convex margin. Median line complete, moderately wide, engraved, joining anterior transverse line. Anterior transverse line narrow, complete. Disk with few scattered punctures at middle, with transverse wrinkles, with isodiametric reticulation at base and laterally, with setigerous punctures situated beside channel. Lateral margin from posterior angle to flange of base slanted with angle of around 40°. Basal channel on flange slightly widened, rugose, basal carina as wide as channel, sharp anteriorly.

Elytron ( Fig. 13 View FIGURES 3–14 ) with disk in lateral view flattened in anterior two thirds, in frontal view slightly convex at middle and moderately convex laterally. Shape of elytra elongate, around two times longer than wide, with lateral margin nearly straight but diverging in anterior third, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus (50-fold). Row of setigerous punctures in lateral channel regular in size, slightly more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining humerus, seventh shortened at humerus, one and two ending free at apex, three and four and five and six joining apically. Intervals slightly convex on disk, convexity more distinct laterally, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven convex up to apex, eight carinate at apex. Surface of all intervals with distinct and regular isodiametric reticulation already traceable at magnification of 40-fold.

Hind wing fully developed.

Lower surface. Proepisternum with isodiametric reticulation. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with slight flattened projection posteriorly at middle, with narrow marginal furrow at middle.

Legs. Protibia with longitudinal reticulation, with wide sulcus dorsally, with three apically slender spines and distinct tubercle, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with one acute and one blunt tubercle with three setae. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front legs slightly widened.

Male genitalia unknown.

Female gonocoxites and epipleurite ( Fig. 111 View FIGURES 110–128 ). Basal gonocoxite inconspicuous, apically with two small setae, apical gonocoxite of moderate size, medially with one very long seta, with three moderately long setae, with two long setae apically. Epipleurite triangle-like, with two setae at middle.

Variation. In the two specimens, the neck constriction is more or less punctured. The anterior transverse line of the pronotum is more or less engraved.

Distribution. Known from two localities in Indonesia, Papua at the Palmer River and from Samabusa near Nabire.

Diagnosis. A slender medium sized fulvous species with peltate pronotum, all intervals of the elytron reticulated, and with six-setose labrum. The most similar species are Clivina dealata brachyptera Darlington, C. hartmanni sp. nov., and C. spatulifigura sp. nov., all of them with isodiametric reticulation on all intervals of the elytron. Out of these, C. opacielytra sp. nov. is the only species with a six-setose and anteriorly pointed labrum. Mainly due to its falciform gonocoxites, C. dealata brachyptera belongs to a different species group, and the reticulation pattern on the elytron is different. In C. hartmanni sp. nov. and in C. spatulifigura sp. nov., the labrum is distinctly excised anteriorly. Moreover, both of them have interval seven of the elytron step-like carinate at apex.

Etymology. The name refers to the elytra (Greek: elytra = ceilings of alae) which are distinctly and completely isodiametrically reticulated giving the surface a dull opaque appearance (Latin: opacus, a, um = dull, like with shadow).