Bryopharsos claviformosum Quate, 1996

Jaume-Schinkel, Santiago, Kilian, Isabel C., Pazmiño-Palomino, Alex & Mengual, Ximo, 2025, Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species, European Journal of Taxonomy 1001 (1), pp. 1-51 : 22

Bryopharsos claviformosum Quate, 1996

Figs 1 , 3C–D

Bryopharsos claviformosum Quate, 1996: 41 . Type locality: Costa Rica, Heredia, Estación Biológica La Selva (INBio).

Bryopharsos claviformosus – Bravo & Araújo 2019: 370 (lapsus calami in identification key).

Diagnosis

Male

Eye bridge with four facet rows; wing 2.5 times as long as wide; ejaculatory apodeme rectangular in ventral view; gonocoxal apodeme with a spine; surstyli with five tenacula. This species is similar to B. amazonensis and B. clavigum but it can be easily differentiated by the number of tenacula (four in B. clavigum , five in B. amazonensis and B. claviformosum ) and the length of the tenacula (equal length in B. amazonensis and B. clavigum , four long and one shorter in B. claviformosum ).

Female

Unknown.

Material examined

ECUADOR – Pichincha • 2 ♂♂; Pedro Vicente Maldonado, near San Pancracio, roadway to Pachijal ; 0.1156° N, 78.9580° E; alt. 750 m; 1–9 Feb. 2022; Isabel Kilian leg.; ZFMK, ZFMK-TIS-2637066, ZFMK-TIS-2637079 • 1 ♂ same data as for preceding; 0.115611° N, 78.95805° E; MECN, ZFMK- TIS-2637137 .

Distribution

Costa Rica ( Quate 1996; Bravo & Araújo 2019) and Ecuador (this publication, new record) ( Fig. 1 ).

DNA barcodes

Three specimens were successfully sequenced: ZFMK-TIS-2637066, ZFMK-TIS-2637079, and ZFMK- TIS-2637137. The maximum intraspecific uncorrected pairwise distance for COI sequences was 0.31% or 2 bp.

Remarks

The original description of the male by Quate (1996) is rather short and incomplete, with some important characters missing. Nonetheless, the drawings and the general description are enough to distinguish the males of this species. In the SEM pictures ( Fig. 3C–D ), the aedeagus seems curved, with a rounded apex and the paramere seems digitiform and slender, while in the original description, the aedeagus looks straight and tapering towards the apex, while the paramere seems broad. We must point out that for our specimens prepared on permanent slides, the perspective affects the perceived shape of the aedeagus and paramere, making the aedeagus look straight and tapering towards the apex, and the paramere looks digitiform.

Figures

Fig. 1. Species distribution map. Species dots do not represent the actual localities; jitter was added for visualization as the species’ real distributions overlap.

Fig. 3. SEM pictures of male terminalia (ventral view). A–B. Bryopharsos amazonensis Bravo & Araújo, 2019. C–D. Bryopharsos claviformosum Quate, 1996. Colors: blue = paramere; cyan = gonocoxal lobes; green = hypandrium; orange = gonostyli; red = aedeagus; yellow = gonocoxites. All scale bars are in millimeters.