Alangium villosum (Blume) Wangerin

de Wilde, W. J. J. O. & Duyfjes, B. E. E., 2017, The species of Alangium section Rhytidandra (Alangiaceae), Blumea 62 (1), pp. 75-83 : 82

11. Alangium villosum (Blume) Wangerin

Alangium villosum (Blume) Wangerin (1906) 61; Bloemb. (1935) 269,f. 3a,f; (1939) 198. ― Marlea villosum (Blume) Kurz (1871) 61. ― Styrax villosum Blume (1826) 671. ― Lectotype here designated: Blume s.n. (L L0009830; iso L 3 sheets L0614166, L0614168, L0614169), Java.

Alangium villosum (Blume) Wangerin var. salaccense (Koord. & Valeton) Bloemb. (1935) 271, nom. illeg. ― Alangium villosum (Blume) Wangerin subsp. salaccense (Koord. & Valeton) Bloemb. (1939) 202, nom. illeg. ― Marlea vitiensis (A.Gray) Benth. var. tomentosa (F.Muell.) Benth. forma salaccensis Koord. & Valeton (1900) 75, nom. illeg.

Alangium villosum ( Blume) Wangerin subsp. tomentosum (F.Muell.) Bloemb. var. javanum Bloemb.(1939) 204,f. 6b, 7a,f. ― Lectotype here designated: Koorders 38191 (L L.2500069;iso BO n.v.), Java, Pasoeroean, Mt Ardjoeno , 1400 m alt., 10 Nov. 1899, fl.

Alangium zollingeri Baill. (1864) 195. ― Type: Zollinger 785Z (holo P P04552612 ), Java.

Alangium villosum (Blume) Wangerin var. tomentosum auct. non (F.Muell.) Bloemb.: Bloemb. (1935) 271, f. 3a, f.

Tree presumably of low stature (but Backer (1965) says 10–20 m tall); twigs (1.5–) 2–4 mm diam, (greyish) brown-hairy, hairs 0.2–0.5(–1) mm long. Leaves: petiole 0.3–1(–1.2) cm long, hairy; lamina drying brown, hairy like the twigs, glabrescent above, ovate-elliptic or (narrowly) elliptic, 6–14(‒19) by 2–5(–7) cm, base little or strongly asymmetric, rounded or short- or long-cuneate, apex acute-acuminate; veins pinnate, 5 or 6 at each side, tertiary venation reticulate, sometimes ± scalariform, domatia absent. Inflorescences solitary peduncled simple or compound cymes, hairy as the twigs, (2–)5–30-flowered; peduncle (3–) 5–15 mm long. Flowers densely hairy like the twigs; pedicel 2–4 mm long; corolla in bud (6‒) 7–8 mm long, base not or little swollen, apex (sub)obtuse; ovary and calyx c. 2 mm long, not ribbed; limb c. 0.5 mm long, at margin c. 2 mm wide, with 5 low triangular lobes c. 0.5 mm long; petals (4–)5(–7), inside (sparingly or towards base strongly) hairy, not connate at base, c. 8 mm long; stamens 5, not adnate to petals, filament c. 3 mm long, hairy, anthers c. 4 mm long, connective glabrous; style 5–6 mm long, glabrous, stigma lobes c. 1 mm long. Fruit 2–7 per infructescence, ripening dark red ( Backer 1965), glabrescent, ovoid or broadly ellipsoid, (8–) 10–15 mm long, smooth or inconspicuously ribbed; calyx remnant minute.

Distribution ― West and East Java.

Habitat & Ecology ― Montane and lower montane forest; (from lowland to) 700–1800 m altitude; flowering from August to December.

Notes ― 1. Specimens form West Java (incl. the type) are rare and scanty and apparently more slender of stature (leaves narrower, inflorescences poorer flowered) as compared with those from East Java. Recent collections are not known .

2. According to Koorders & Valeton (1900) the bark and leaves are very bitter.