Elymus nevskii Tzvelev

Alieva, Kumush B., Peng, Yilong, Usupbaev, Adilet, Tojibaev, Komiljon Sh., Yusupov, Ziyoviddin, Ergashov, Ibrokhimjon, Azimova, Dilnoza & Jiang, Zhilin, 2025, Synopsis of the genus Elymus (Poaceae) in Uzbekistan (Middle Asia) with a description of Elymus uzbekistanicus a new species from Turkestan Mts, PhytoKeys 257, pp. 9-50 : 9-50

6. Elymus nevskii Tzvelev in Spisok Rast. Gerb. Fl. S. S. S. R. Bot. Inst. Vsesojuzn. Akad. Nauk 18: 29 (1970)

= Agropyron dentatum Hook. f. in Fl. Brit. India 7: 370 (1896). Type. India. Kashmir, alt. 9–12,000 ft., Jacquemont, Thomson.

= Agropyron ugamicum Drobow in A. I. Vvedensky & al., Key Fl. Tashkent 1: 41 (1923). Type. Uzbekistan. Western Tian Shan. “ Distr. Tashkent, near the Ugam River, ” 1921, n ° 1313, Uranov. ≡ Semeiostachys ugamica (Drobow) Drobow in Fl. Uzbekistan. 1: 284 (1941). ≡ Roegneria ugamica (Drobow) Nevski in Trudy Sredne-Aziatsk. Gosud. Univ., Ser. 8 b, Bot. 17: 69 (1934).

Type.

Tajikistan • Leninabad region, southern slope of the Zeravshan range, in the basin of the Yagnob River, between the tributaries Golkrod and Yamansu , on a subalpine meadow, at an altitude of 2900 m. 18 VII 1935, V. Nikitin [ K 003372662 ] .

Description.

Stems 50–120 cm, glabrous, smooth. Leaves 7–11 mm wide, flat, abaxially smooth and glabrous, adaxially rough with scattered hairs. Lowest part of sheath hairy. Ligule short. Spikes straight, 7–13 cm. Spikelets 20–30 mm, green or purple. Internodes of spike rough only along two lateral ribs. Glumes broadly lanceolate, margins membranous, quickly acuminate to bristle-like apex to 2 mm. Lemma abaxially nearly or entirely densely spiny or hairy, with awn to 7 mm. Palea linear, nearly equal to lemma. Anthers 1.8–3 mm (Fig. 8 ). 2 n = 28 ( Tzvelev and Probatova 2019).

Phenology.

Flowering and fruiting: June-August.

Habitat.

Middle, subalpine, and alpine mountain belts, among shrubs (polydominant, mesophytic deciduous), meadows (floodplain, mesophytic), petrophytic communities (rocky-gravelly scree, cliffs), and gravel, 800–3200 m.

General distribution.

China, Pakistan, Russia, western Himalayas, Middle Asia (Tarbagatai, Junggar Alatau, Tian Shan, Aral-Caspian Lowlands, Pre-Balkhash Deserts, Pamir-Alay: Gissar, Darvaz, and Pamir ranges): Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan.

Distribution in Uzbekistan.

Jizzakh, Kashkadarya, Namangan, Surkhandarya, and Tashkent regions (Fig. 3 F ). I- 1 Western Tian Shan district. I- 1 - a Ugam-Pskem region. UGAM RANGE (Syr-Darya Region, Tashkent District. Valley of the Ugam River. 06.1921, Maksimov s. n. [MW 0808955]; Valley of the Pskem River. Babajan Sai (left tributary of the Pskem River). On turf-covered slopes, 26. 07. 1941, Momotov 84 [TASH 055307!]; Syr Darya Region, Tashkent District. Valley of the Ugam River, Maksimov s. n. [TASH 055308!]; Bostandyk, on rocky slopes along the ridge of the Ugam Range in the upper reaches of the Navali-Sai gorge, at 2800–3000 m, 31. 07. 1953, Pavlov 467 [MW 0808951, MW 0808959]; Oygaing, 03. 08. 2023, Turdiev, Aliyeva O _ 16 _ A [TASH 054576!]); PSKEM RANGE (Bostandyk, on rocky scree beneath a large snow patch at the head of Aksar-Sai, 24. 07. 1949, Pavlov 158 [MW 0808948], 28. 07. 1949, Pavlov 244 [MW 0808949, MW 0808950]; Upper reaches of the Oygaining River. Gentle slopes of the right bank of Tunduk-Sai, 20. 08. 1956, Tsukervanik 1572 [TASH 055292!], Granitov 1567 [TASH 055296!]). I- 1 - b Western Chatkal region. CHATKAL RANGE (Chatkal Mountain-Forest Reserve. Maidan-Tal section, Tashkent-Say, southern slope at an altitude of 1500–1700 m, 10.10., Savich s. n. [TASH 046203!, TASH 046204!]; Vicinity of the Chimgan Botanical Station. Gorge of the Chimganka River, on the stream bank near the waterfall, 26. 08. 1928, Gomolitsky 477 [TASH 055301!]; Myn-Dzhilke tract. Upper reaches of the Nurek-Ata River. Southern gravelly slopes (does not fit the description, as there is no awn on lemma), 17. 07. 1936, Korotkova, Titov 1430 [TASH 047501!], 1446 [TASH 047502!]; Basin of the Angren River. Rocky slope below the Davan-Sai Pass, 15. 08. 1937, Zakirov s. n. [SAMDU]; Basin of the Chatkal River. Mazar River Valley. Eastern slope of the left bank of the Mazar River, belt of creeping form juniper, 08. 08. 1938, Pyataeva, Momotov 604 [TASH 055304!]; Tashkent Alatau Mountains. Basin of the Bash-Kyzyl-Say River. Menora-Say, 02. 08. 1953, Butkov, Tsukervanik 1203 [TASH 046343!]; New section of Reserve, Chatkal Mountains, 70 km E of Tashkent, 314209 [USDA _ NPGS 3417113], 314200 [USDA _ NPGS 3417106], 314211 [USDA _ NPGS 3417103]); I- 1 - c Arashan region. CHATKAL RANGE (Angren Expedition. Right bank of the Angren River; warm springs of Arasan, granite, 14. 08. 1924, Korovin 2736 [TASH 055326], 16. 08. 1924, Korovin 2737 [TASH 055327]; Basin of the Angren River. Upper reaches of the Angren River, above the warm springs of Arashan, along the shore of the lake, at an altitude of 1750 m. 28. 07. 1938, Pyataeva, Momotov 347 [TASH 055329!, TASH 055330!]; Angren Plateau. Southern slopes. Near Arasan Lake. Northeastern slope. Alpine vegetation belt, 19. 08. 1939, Kudryashev 1237 [TASH 055331!]; Arashan Lake, on the slopes, 24. 07. 1940, Korotkova 231 [TASH 047493!]); I- 1 - d Kurama region. CHATKAL RANGE (Basin of the Angren River. Upper reaches of the Angren River, above the warm springs of Arashan, sand deposits on the southeast slope of the Babai-Tagh ridge, site near the water, 28. 07. 1938, Pyataeva, Momotov 316 [TASH 046200!]; Tashkent Alatau. Southern gravelly slope of Mountain Kyzyl-nura, altitude 3200 m, 11. 09. 1939, Gomolitsky 77 [TASH 055309!]; Basin of the Angren River. In the upper reaches of the Chet-su, on gravelly slopes, 27. 07. 1940, Korotkova 315 [TASH 055335!]; Valley of the Angren River. Sai Ayri. Tugai, 17. 08. 1940, Usmanov 1187 [TASH 055386!]); KURAMA RANGE (Kishvash Sai, upper reaches of Lashkerek, above the juniper zone. Rocky slope, 12. 07. 1939, Zakirov s. n. [SAMDU]; Basin of the Angren River. Dawan-sai, rocky slope below the pass, 15. 08. 1937, Zakirov s. n. [TASH 055328!]; Northern slopes of the upper reaches of Nizbash-sai. Eastern slope with subalpine vegetation, 06. 08. 1939, Kudryashev 1083 [TASH 055336!, TASH 055337!]; Northern slope of Say Saracha, western slope of the Juniper forest, 13. 07. 1940, Usmanov 1307 [TASH 055390!]; Northern slope of Say Kum-Kul. Tugay vegetation, 10. 08. 1940, Usmanov 1210 [TASH 055387!]; Northern slopes of the Kurama Range, Abyazsay, 4 th brigade of the breeding sovkhoz, h = 1950 m, 11. 09. 1956, Kamalov 136 [TASH 055389!], h = 1900 m, juniper zone (prostrate form), of the breeding farm, 11. 09. 1956, Kamalov, Vernik, Nabiev, Tsukervanik 137 [TASH 055388!], Stud farm, rocky gravelly slope, 2150 m, 17. 09. 1956, Kamalov 316 [TASH 055385!]). I- 5 Kuhistan district. I- 5 - a North Turkestan region. TURKESTAN RANGE (Zaamin Forestry, Gorge of the Kul-su River, 23. 07. 1926, Popov, Androsov 116 [TASH 055323!], 117 [TASH 055324!]; Gurlash River Gorge, on the left bank of the Gurlash River, opposite Chutka-Say, 25. 07. 1926, Popov, Androsov 144 [TASH 055325!]; Zaamin District. Headwaters of the Chandyir Rivers, near the pass. Absolute altitude 2700–3000 m, 14. 09. 1932, Titov, Eliseeva 600 [TASH 055322!]; Pamiro-Alay. Basin of Guralash River. Cone of the Mechetyly-Say River, 27. 07. 1934, Zakrzhevskiy 491 [TASH 055316!, TASH 055319!]; Northern slopes of the Turkestan Range, headwaters of the Sanzar River. Guralash-Say Nature Reserve. At the head of Guralash, on the northern slopes of Langar-Say, 14. 08. 1937, Korotkova, Vasil’kovskaya 1039 [TASH 055341!], 17. 08. 1937, Korotkova, Vasil’kovskaya 1073 [TASH 055320!]; along the ridge of Angerly-Say, opposite Shibarly-Say. Dry subalpine meadow, 30. 08. 1947, Nazarenko s. n. [TASH 055237!, TASH 055238!]; 31. 08. 1947, Nazarenko 634 [TASH 055235!]); I- 5 - b Malguzar region. MALGUZAR RANGE (Northern slopes of the Turkestan Range. 4–5 km from the village of Besh-Kubu towards the Zaamin mountains, 22. 05. 1937, Korotkova, Vasil’kovskaya 989 [TASH 055338!]). I- 6 Western Hissar district. I- 6 - a Kashkadarya region. HISSAR RANGE (Mountain Hazret-Sultan. Aksu River. Elevation 2800 m, 17. 07. 1933, Gordienko, Chilinyn 277 [MW 0808944]). I- 7 Hissar-Darvaz district. I- 7 - a Sangardak-Tupalang region. HISSAR RANGE (Pamiro-Alay. Upper reaches of the Yagly-Khocha [Yangiklik] River, elevation 3200 m. Description N 82, 14. 08. 1941, Gromakov 722 [TASH 055318!]; Basin of the Tupolang River. Upper reaches of the Khovar River. Eastern slope in the alpine zone, 09. 09. 1947, Pyatayeva 603 [TASH 040959!]; Above Dara-Say at the pass. Subalpine and alpine zones, 17. 07. 1948, Pyatayeva 1084! [TASH 055317!]).

Figures

Figure 3. Distribution maps of Elymus species in Uzbekistan A E. caninus B E. lachnophyllus C E. fedtschenkoi D E. longearistatus E E. macrochaetus F E. nevskii G E. praeruptus H E. transhyrcanus I E. tschimganicus J E. tianschanigenus.

Figure 8. Elymus nevskii. a, spike b spikelet c glumes d lemmas and paleas e stem f node g ligule (https://doi.org/10.13140/RG.2.2.22526.60481).