Anomala sabatinellii Zhao & Pham, 2023

Zhao, Ming-Zhi, Pham, Phu Van & -, P. R .., 2023, Two new species of the genusAnomala Samouelle from Vietnam and Laos, with a checklist of Vietnamese Anomala (Coleoptera: Scarabaeidae: Rutelinae), Faunitaxys (New York, N. Y.) 11 (55), pp. 1-11 : 1-2

Anomala sabatinellii Zhao & Pham sp. nov.

( Fig. 1- 5, 9- 11)

ZooBank:http://zoobank.org/ F2B0497E-46ED-42AB-97B7-7D5079520803

Holotype ♂ ( MHNG), “ VIETNAM, Gia Lai Prov., Kon Ka Kinh , 15.VI.2022, local collector leg.”

Paratypes (18 ♂ & 12 ♀)

– 3 ♂ & 1 ♀ ( CGS) , 3♂ ( CZMZ) same data as holotype .

– 4 ♂ ( IEBR), “ VIETNAM, Gia Lai Pro., Kon Ka Kinh NP, close to

Reviewer:

guest house, light trap, 26.iv.2022, P. V. Pham leg.”

– 1 ♂ & 1 ♀ ( CCZ), “ VIETNAM, Kon Tum, Đắk Tô , iv. 2020, local collector leg.”

– 1 ♂ ( CCZ), “ Qu ảng Sơn, Đăk Nông, Vietnam. May, 2018, local collector leg.”

– 2 ♂ ( CCZ), “ LAOS, Attapeau Prov. , Annam Highlands Mts. , Dong Amphan NBCA, ca. 1160 m, NONG FA (crater lake) env., 15°05.9′ N, 107°25.6′ E, St. Jakl lgt. 30.iv-4.v.2010 ” .

– 1 ♂ & 2 ♀ ( CCZ) , 1 ♂ & 3 ♀ ( CSJ), “ LAOS, Attapeau Prov. , Annam Highlands Mts. , Dong Amphan NBCA, ca. 1160 m, NONG FA (crater lake) env. 15°05.9′ N, 107°25.6′ E 30.iv-6.v.2010 St. Jakl lgt.”

– 3 ♀ ( NMPC) , 1 ♀ ( CCZ), “ LAOS, ATTAPEU prov., Annam Highlands Mts., Dong Amphan NBCA, ca. 1160 m, NONG FA [crater lake], env., 15°05.9′ N, 107°25.6′ E, Jiří Hájek leg. 30.iv-6.v.2010 ” .

– 2 ♂ & 1 ♀ ( CMF), “S.E. Laos, Dakchung 1300 m alt., 1-5.I.2011 ” (in Sekong Province) .

Description of the holotype male ( Fig. 1 -3, 9 - 11)

General appearance. – Body shape elongated ovoid, strongly convex. Generally yellowish brown, with greenish sheen and iridescence; frons and vertex, two elongated large patches on pronotum, suture of elytra, anterior margin of propygidium, two large spots on each anterior angle of pygidium, mesosternum, joints and apices of all tibiae, as well as all tarsi green to dark green; borders of each abdominal ventrite and surroundings of disc of metasternum indefinitely green; antennal club black; setae yellowish brown, spiniform setae dark brown.

Head. – Clypeus subtrapezoidal, anterior corner obsolete and widely arched; anterior margin feebly reflexed; surface with very dense and coarse punctures, somewhat rugose basally. Frontoclypeal suture distinct. Frons with very dense and coarse punctures, gradually finer posteriad, vertex with transverse and small punctures. Antennal club almost equally as long as antennomeres 1–6 combined. Inner margin of eye with several moderately long to long setae.

Pronotum. – Sides gently arched and convergent anteriad, anterior half inconspicuously straight. Anterior angle acute and protruding, posterior angle rounded. Basal marginal line broadly interrupted before scutellum, other marginal lines complete. Disc with transverse, dense and large punctures, punctures horizontally elongated and coalescent at lateral portions. Lateral margin with a few moderately long to long setae.

Scutellum . – Almost triangular, lateral margin arched. Glabrous, disc punctate as disc of pronotum.

Elytra . – Intervals strongly convex forming costae, subsutural interstices widest with two costae, other intervals each with one costa, almost equal in width; strial punctures very dense, transverse and coalescent, forming numerous transverse striolation across each two adjacent intervals; tip of the convex intervals smooth, sparsely bearing small punctures. Humeral umbone and apical protuberances moderately prominent. Lateral carina distinct at basal three fourth. Epipleura with a row of sparse and short setae. Marginal membrane complete and broad.

Propygidium . – Surface with dense and transverse striolation. With a transverse row of sparse short setae proximally.

Pygidium . – Surface strongly tumid proximally. Surface with dense and transverse striolation encircling the greatest tumidity. With sparse and irregularly-distributed long setae proximally, and a transverse row of long setae at posterior margin.

Ventralthoracicsurface. – Ventral pro- and mesothoracic surface with very dense, longitudinal or oblique, and frequently coalescent punctures, sparsely bearing short setae. Ventral metathoracic surface with very dense punctures, punctures on medial portions moderately dense and small, then gradually larger and dense laterad, all punctures becoming transverse and coalescent at lateral portions, dense setae being short to moderately long.

Abdominal ventrites. – Ventrites not carinate laterally. Punctate as in metasternum, but the punctures on medial portions transverse. Ventrites 2–5 each with a transverse but sparse row of short or moderately long setae, the row of ventrite 6 present at posterior margin.

Legs. – Protibia bidentate, both teeth distinct; apical tooth extending to level of halfway of protarsomere 2. Inner spur inserted shortly behind the level of the proximal tooth. Inner protarsal claw and outer mesotarsal claw split apically; lower branch of inner protarsal claw convex at lower margin basally, forming a notch internobasally, shorter anddistinctly wider than the sharp upper branch; lower branch of outer mesotarsal claw longer and distinctly wider than upper branch. Each tarsomere 5 with an internomedial denticle, that of mesotarsomere 5 small. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with longitudinal rows of short setae, two regular rows next to dorsal margin, a third row situated behind middle and bearing spiniform setae, a row at ventral edge with fine setae. Metafemoral surface with a row of sparse, short and spiniform setae behind middle, ventral edge with a few fine setae.

Male genitalia. – As Fig. 9 -11. Parameres weakly asymmetric and finely setose; ventral plate strongly hooked proximally.

Male paratypes ( Fig. 4). – External featuresandshape of parameres consistent. A male paratype ( Fig. 4) has generally green dorsal surface and protibia, only clypeus and lateral portions of pronotum are yellowish brown.

Female paratypes ( Fig. 5). – Generally similar to male. Antennal club slightly shorter than antennomeres 1–6 combined. Pygidium almost triangular in shape and weakly tumid. Setae on ventral surface distinctly denser and longer.Apical protibial tooth abraded, proximal tooth broadly rounded; inner spur inserted behind halfway of protibia; internomedial protuberance of protarsomere 5 small; upper branch of inner protarsal claw slightly longer than the lower one, a denticle devoid on lower branch, thus not forming a notch internobasally; metatibia wider than in male. Ultimate abdominal ventrite almost one time longer than in male.

Measurement. – Body length: 14.8-17.3 mm for male and 15.7-18.5 mm for female, greatest width: 8.1-9.5 mm for male and 9.0- 10.4 mm for female.

Differential diagnosis. – Anomala sabatinellii Zhao & Pham sp. nov. is closely allied to A.immeliorata Zorn, Kobayashi & Wada, 2017 and A. confrater Zorn, Kobayashi & Wada, 2017 , both described from Vietnam.The rightparamere of A.confrater bearsastrongdenticleatits halfway and ventral plate simply bends downward, which allows an easy separation. In A. immeliorata , the parameres are spike-liked in apical half, while the parameres are strongly compressed and bear a ridge ventrally in A.sabatinellii . What’s more, the ventral plate laterally and proximally forms a small hook (gap ca. 0.5 mm) in A.sabatinellii but a large hook (gap ca. 1.5 mm) in A. immeliorata .

Etymology. – This species is named after Dr. Guido Sabatinelli, who generously provided a series of this taxon for study.

Distribution. – Vietnam: Đắk Nông, Gia Lai, Kon Tum; Laos: Attapeu, Sekong.