Haustrum? pyrulatum (Hoernes & Auinger, 1882)
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publication ID |
https://doi.org/10.11646/zootaxa.5572.1.1 |
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publication LSID |
lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242 |
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persistent identifier |
https://treatment.plazi.org/id/F73C87F9-FFE1-8053-FF50-AF4CF9ABB9EB |
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treatment provided by |
Plazi |
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scientific name |
Haustrum? pyrulatum (Hoernes & Auinger, 1882) |
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Haustrum? pyrulatum (Hoernes & Auinger, 1882)
Figs 1 View FIGURE 1 , 2A View FIGURE 2 1 –A View FIGURE 1 2 View FIGURE 2
* Purpura (Polytropa) pyrulata nov. form.—Hoernes & Auinger 1882: 152, pl. 16, fig. 9.
Type material. Holotype: NHMW 1859 View Materials /0045/0590, SL: 32.5 mm, MD: 19.6 mm, Lăpugiu de Sus ( Romania),
illustrated in Hoernes & Auinger (1882: pl. 16, fig. 9), Figs 2A View FIGURE 2 1 –A View FIGURE 1 2 View FIGURE 2 .
Revised description. Medium-sized, ovate shell; apical angle 63°. Protoconch unknown. Teleoconch of at least four whorls. Suture distinctly incised, linear. Early teleoconch whorls moderately convex with rounded shoulder. Spiral sculpture of very broad, strap-like, rounded IP, P1 and P2 separated by narrower interspaces on antepenultimate and penultimate whorls. No axial sculpture except for narrow growth lines forming cancellate pattern in spiral interspaces. Last whorl ovate with convex periphery, moderately constricted below. Very broad,
band-like IP, P1–P6, ADP and narrow s1 with deep, narrow interspaces. P1 only slightly more prominent. Fasciole moderately swollen; narrow pseudoumbilicus covered by columellar callus. Aperture wide, ovate; outer lip not thickened (margin missing) with blunt D1–D5 placed some distance behind peristome. Anal canal wide, poorly delimited. Siphonal canal moderately short, wide, open, slightly bent to the left. Columella broadly excavated, smooth. Columellar callus thin, adherent, poorly delimited from base.
Discussion. Extent Haustrum species occur in the south Pacific. Therefore, its occurrence in the Circum-Mediterranean Neogene would be surprising but Haustrum pyrulatum (Hoernes & Auinger, 1882) is strikingly similar to the extant Haustrum lacunosum ( Bruguière, 1789) from New Zealand ( Figs 2B View FIGURE 2 1 –B View FIGURE 1 2 View FIGURE 2 ). Both species differ from each other only by the narrow s 1 in Haustrum pyrulatum .
Haustrum pyrulatum was originally placed in Polytropa Swainson, 1840 by Hoernes &Auinger (1885), which is a junior synonym of Nucella Röding, 1798 [type species Nucella lapillus ( Linnaeus, 1758) ], subsequent designation by Stewart (1927: 386); present-day, northern Atlantic]. Indeed, Haustrum pyrulatum is also reminiscent of the Ocenebrinae genus Nucella . Nucella species differ from Haustrum pyrulatum in their narrower primary cords and the more prominent secondary cords, which are often equal to the primary cords in strenght. Moreover, Nucella is a cool-temperate genus, which originated during the late Middle Miocene in the Pacific and spread to the Atlantic around 6.2 my ago ( Marko et al. 2014). Therefore, a close relation of the Paratethyan species with Nucella is unlikely. Spinucella Vermeij, 1993 [type species Purpura tetragona J. de C. Sowerby, 1825, original designation by Vermeij (1993: 20); Pliocene, North Sea] occurred also in the warmer Proto-Mediterranean waters during the Miocene and Pliocene (e.g., Spinucella monocanthos ( Brocchi, 1814) , S. cancellata ( Bellardi, 1882) , S. depressa ( Bronn, 1831) but these species usullay have a more or less shouldered last whorl and differ from Haustrum in their narrower spiral cords and the presence of axial ribs (see Bellardi 1882: pl. 12, figs 1–3). Although we note the problem of a Paratethyan distribution of an otherwise south Pacific genus, we tentatively place this species in Haustrum with questionmark. However, it might represent a striking example of convergence.
Paleoenvironment. Unknown. Both, shallow water and deep water species occur at Lăpugiu de Sus ( Romania) (own data M.H.).
Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1882).
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