Ocenebra vindobonensis (Hörnes, 1853)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 60-62

Ocenebra vindobonensis (Hörnes, 1853)

Figs 13 , 27A–E

* Murex Vindobonensis Hörnes —Hörnes 1853: 252 (pars), pl. 25, figs 17, 20.

Murex (Phyllonotus) Vindobonensis M. Hoern. —Hoernes & Auinger 1885: 213 (pars).

P [urpura]. (T [ritonalia].) vindobonensis (Hörn.) — Sieber 1958: 146.

[ Ocenebra ] vindobonensis (Hoernes, 1853— Vokes 1971: 116.

Ocenebra vindobonensis (Hoernes, 1853) — Stojaspal 1978: 335.

Hexaplex (Phyllonotus) vindobonensis (M. Hoernes, 1856) — Švagrovský 1982: 385, pl. 1, figs 1–3.

Purpura (Tritonalia) vindobonensis (Hörnes, 1856 [sic])— Bałuk 1995: 227, pl. 30, fig. 5.

Purpura (Tritonalia) vindobonensis (Hörnes, 1856 [sic])—Harzhauser 2002: 97, pl. 6, figs 13–14.

Purpura (Tritonalia) vindobonensis (Hörnes, 1856 [sic])— Bałuk 2006: 209, pl. 13, figs 4–8.

Ocenebra vindobonensis vindobonensis (Hörnes, 1853) — Kovács et al. 2018: 123, figs 7I–J.

Ocenebra vindobonensis vindobonensis (Hörnes, 1853) — Kovács 2019: 130.

Ocenebra vindobonensis vindobonensis (Hörnes, 1853) — Kovács 2020: 457, pl. 3, figs 5–7.

non Murex Vindobonensis Hörnes —Pereira da Costa 1867: 167, pl. 20, figs 4a–b [= Ocenebra mirmigus group]. non Murex (Phyllonotus) Vindobonensis M. Hoern. —Hoernes & Auinger 1885: 213, pl. 25, fig. 2 [= Ocenebra mirmigus (De

Gregorio, 1885a)]. non Tritonalia vindobonensis (Hörnes) var.— Kókay 1966: 57, pl. 8, fig. 6 [= Ocenebra mirmigus (De Gregorio, 1885a) ]. non Ocenebra vindobonensis (Hörnes, 1856 [sic])— Goret & Pons 2013: 61, text-fig 5/2, pl. 4, fig. 2 [? = Ocenebra ligeriana

( Tournouër, 1875)]. non Ocenebra vindobonensis vindobonensis (Hörnes, 1853) —Landau et al. 2016: 224, pl. 1, fig. 4 [= Ocenebra breitenbergeri

sp. nov.].

Type material. Lectotype designated by Švagrovský (1982: 385): NHMW 1851 /0026/0028d, SL: 29.2 mm, MD: 16.1 mm, Grund ( Austria), illustrated in Hörnes (1853: pl. 25, fig. 17), Figs 27A 1 –A 3 . Paralectotypes: NHMW 1851 /0026/0028e, SL: 35.9 mm, MD: 21.9 mm, Grund ( Austria), Figs 27B 1 –B 3 .

Illustrated material. NHMW 1851/0026/0028f, SL: 32.4 mm, MD: 18.5 mm, Grund ( Austria), Figs 27C 1 –C 3 . NHMW 1851/0026/0028g, SL: 31.4 mm, MD: 17.9 mm, Grund ( Austria), Figs 27D 1 –D 2 . NHMW 1851/0026/0028, SL: 35.9 mm, MD: 22.0 mm, Grund ( Austria), Figs 27E 1 –E 2 .

Additional material. NHMW 1846 /0037/0230, SL: 28.3 mm, MD: 16.4 mm, Enzesfeld ( Austria), illustrated in Hörnes (1853: pl. 25, fig. 20); 4 spec. , NHMW 1868 /0001/0217, Grund ( Austria); 32 spec. , NHMW 1868 /0001/0216, Grund ( Austria); 3 spec. , NHMW 1847 /0037/0062, Gainfarn ( Austria); 4 spec. , NHMW 1857 /0002/0050, Bad Vöslau ( Austria); 5 spec. , NHMW 1870 /0037/0017, Ritzing ( Austria); 3 spec. , NHMW 1860 /0001/0199, Boršov (= Porstendorf) ( Czech Republic); 13 spec. , NHMW1866 /0040/0286, Lăpugiu de Sus ( Romania) .

Revised description. Medium-sized, broad fusiform, trivaricate shell with conical spire; apical angle ~55–65°. Protoconch unknown.Teleoconch of up to five whorls.Suture weakly incised, shallowly undulating. Early teleoconch whorls convex with faint shoulder. Axial sculpture of eight to nine prominent ribs, overrun by delicate adis, IP and abis and prominent P1–P2 on antepenultimate and penultimate whorls. Later teleoconch whorls with broad, steep subsutural ramp and marked shoulder. Last whorl attaining ~80% of total height; three raised, weakly foliaceous varices and three weaker intervarical ribs. Subsutural ramp with subequal adis, IP and abis and only slightly weaker tertiary cords. P1 strong, P2 atrophied, P3 of medium strength, P4, P5 roughly equal to P1, P6 atrophied, forming spines of medium length on varices; ADP, MP weak. Secondary cords weaker, s3 and s4 occasionally forming short spines on varices; s1, s2, s5 and s6 occasional and weaker. Fasciole prominent, swollen, delimiting very broad pseudoumbilicus. Aperture ovate; outer lip with crenulate edge, prominent D1–D5, often bifid (particularly D2 to D4), forming close-set, paired denticles close behind peristome. Anal canal broad, poorly delimited; siphonal canal relatively short, narrow, fused, bent to the right. Columella straight, smooth, slightly twisted at siphonal canal. Columellar callus forming broad rim, sharply delimited from base, erect at siphonal canal, adherent in parietal area.

Discussion. Vokes (1971: 65) placed Murex vindobonensis ligeriana Tournouër, 1875 in Ocenebra , which is adopted herein also for Murex vindobonensis Hörnes, 1853 . Ocenebra vindobonensis (Hörnes, 1853) differs from the extant Ocenebra erinaceus ( Linnaeus, 1758) in its conical spire and bifid denticles in the outer lip. Labial denticles in O. erinaceus when present are always weak. When Tournouër (1875) described Murex vindobonensis var. ligeriana , he illustrated (pl. 5, fig. 3) a specimen bearing spiny P1 to P5, lacking P6, ADP and MP and with a high spire bearing P1 and P2. Glibert (1952a, pl. 7, fig. 5a–b) illustrated a specimen with very similar characters under the name Tritonalia vindobonensis var. ligeriana , but he illustrated two other specimens (pl. 7, fig 5c–d) devoid of spines. We agree with Glibert (1952a) that these two morphotypes (one spiny and one devoid of spines) represent variations of a single species. The specimen from the Burdigalian/Langhian (Early/Middle Miocene) of Montpeyroux ( France), described as Ocenebra vindobonensis by Goret & Pons (2013, pl. 4, fig. 2) resembles the morphotype devoid of spines of O. ligeriana figured by Glibert (1952a) in its higher spire, in its reduced spiral sculpture on the subsutural ramp, in its shorter spines on the terminal varix and in its simple denticles in the outer lip. The same features distinguish the specimens from the Serravallian of the Karaman Basin ( Turkey), described herein as Ocenebra breitenbergeri sp. nov. from Ocenebra vindobonensis . Finally, O. vindobonensis differs from O. ligeriana by having a lower spire, spiral sculpture on the subsutural ramp (including adis, IP, abis), and a reduced P2.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Kleinebersdorf, Stetten ( Austria) (Harzhauser 2002). Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1995); North Alpine-Carpathian Foreland Basin: Grund ( Austria); Vienna Basin: Bad Vöslau, Enzesfeld ( Austria) (Hoernes & Auinger 1885), Boršov (= Porstendorf), Mikulov ( Czech Republic) (Hoernes & Auinger 1885), Borský Mikuláš ( Slovakia) ( Švagrovský 1982); Danube Basin: Smolenice, ( Slovakia) ( Švagrovský 1982); Oberpullendorf Basin: Ritzing ( Austria) (Hoernes & Auinger 1885); Danube Basin: Smolenice ( Slovakia) ( Švagrovský 1982); Pannonian Basin: Letkés, Hosszúhetény ( Hungary) ( Kovács et al. 2018).

Not in Paratethys:

Figures

FIGURE 1. Cord diagrams and apertural features for Haustrinae [Haustrum? pyrulatum (Hoernes & Auinger, 1882)] and Muricinae [Bolinus gapus (De Gregorio, 1885a), Bolinus partschi (Hörnes, 1853), Chicoreus (Triplex) aquitanicus (Grateloup, 1833), Chicoreus (Triplex) hirmetzli (Kovács, 2018), Chicoreus (Triplex) notatus (Eichwald, 1830). Hexaplex (Trunculariopsis) trunculoides (Pusch, 1837), Hexaplex (Trunculariopsis) esplus (De Gregorio, 1885a), Hexaplex(Trunculariopsis) subasperrimus (d’Orbigny, 1852), Promurex spinicosta (Bronn, 1831), Exquisitor graniferus (Michelotti, 1841)].

FIGURE 2. A1–A2. Haustrum? pyrulatum (Hoernes & Auinger, 1882), holotype, NHMW 1859/0045/0590, Lăpugiu de Sus (Romania). B1–B2. Haustrum lacunosum (Bruguière, 1789), MNHN-IM-2000-904, lectotype, New Zealand, present-day, photo: Manuel Caballer, 2014 RECOLNAT project ANR-11-INBS-0004.

FIGURE 3. Bolinus gapus (De Gregorio, 1885a). A1–A3. NHMW 1847/0046/0030, Szob (Hungary). B1–B3. paralectotype, NHMW 1860/0015/1270, Grund (Austria). C1–C3. NHMW1869/0001/0280, Grund (Austria). D1–D3. lectotype, NHMW 1856/0050/0122, Gainfarn (Austria).

FIGURE 13. Cord diagrams and apertural features of Ocenebrinae [Ceratostoma? steiningeri sp. nov., Gracilipurpura polonica (Bałuk, 1995), Heteropurpura catosus (De Gregorio, 1885a), Kestocenebra subaustriaca (Stojaspal, 1978), Kestocenebra vermeiji sp. nov., Kestocenebra? globosa (Stojaspal, 1978), Ocenebra ergnapus (De Gregorio, 1885a), Ocenebra haidingeri (Hörnes, 1853), Ocenebra holubicensis (Friedberg, 1912), Ocenebra littoralis nov. nom., Ocenebra mirmigus (De Gregorio, 1885a), Ocenebra scorpio sp. nov., Ocenebra valdemari (Bałuk, 2006), Ocenebra vindobonensis (Hörnes, 1853), Ocenebra zboroviensis (Friedberg, 1912)].

FIGURE 27. Ocenebra vindobonensis (Hörnes, 1853). A1–A3. Lectotype, NHMW 1851/0026/0028d, Grund (Austria). B1– B3. Paralectotype, NHMW 1851/0026/0028e,, Grund (Austria). C1–C3. NHMW 1851/0026/0028f, Grund (Austria). D1–D2. NHMW 1851/0026/0028g, Grund (Austria). E1–E2. NHMW 1851/0026/0028, Grund (Austria).