Ocenebra ergnapus (De Gregorio, 1885a)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 49-50

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FFD6-8068-FF50-A9ECF9D2BFAC
treatment provided by	Plazi
scientific name	Ocenebra ergnapus (De Gregorio, 1885a)
status

Treatment

Figs 13 View FIGURE 13 , 20A–E View FIGURE 20

Murex craticulatus Brocc. View in CoL —Hörnes 1853: 234 (pars), pl. 24, figs 9, 11 [non Brocchi, 1814, = Gracilipurpura craticuloides ( Vokes, 1964) View in CoL ].

* [ Murex craticulatus View in CoL ] F. a ergnapus De Greg. —De Gregorio 1885a: 245 [pro Murex craticulatus sensu Hörnes 1853 View in CoL : pl. 24, fig. 9].

[ Murex craticulatus ] F. a perisus De Greg.—De Gregorio 1885a: 244 [pro Murex craticulatus sensu Hörnes 1853: 239 , pl. 24, fig. 11].

[ Murex craticulatus ] F. a erpis De Greg.—De Gregorio 1885a: 245 [pro Murex craticulatus sensu Hörnes 1853: 239 , pl. 24, fig. 11].

Murex ( Occenebra [sic]) Boeckhi nov. form.—Hoernes & Auinger 1885: 221, pl. 27, fig. 3.

Tritonalia boeckhi H.-Au.— Strausz 1954: 24, pl. 5, fig. 122.

H [adriania]. boeckhi (R. Hörn., et Au.)— Sieber 1958: 146.

Hadriania boeckhi Hoernes & Auinger, 1880 [sic]— Strausz 1966: 276, pl. 56, figs 13–14.

[ Ocenebra View in CoL ] boeckhi Hörnes & Auinger, 1885 — Vokes 1971: 24.

[ Ocenebra View in CoL ] ergnapus De Gregorio, 1885 — Vokes 1971: 45.

Ocenebra (Ocenebra) boeckhi (Hörnes—Auinger, 1885)— Stojaspal 1978: 337, pl. 5, fig. 2.

Hexaplex (Muricanthus) sandbergeri (Hörnes, 1856) — Mikuž 2009: 26, pl. 8, fig. 107 [non Pterynotus (Pterymarchia) sandbergeri (Hörnes, 1856) View in CoL ].

Ocinebrina boeckhi (Hoernes et Auinger) — Kovács & Vicián 2024: 15, figs 33–34.

non Murex View in CoL ( Hadriania View in CoL )? sp. cf. craticulatus View in CoL L.— Sokolov 1899: 34, pl. 4, figs 14–17.

non Hadriania cf. craticulata View in CoL — Bidzinashvili 1975: 106, fig. 6.

non Hadriania boeckhi (R. Hoernes et Auinger, 1885) — Iljina 1993: 100, pl. 13, figs 10–11.

non Ocinebrina boeckhi (Hoernes & Auinger, 1885) — Kovács et al. 2018: 122, figs 5W–X [= Pterynotus (Pterymarchia) sp.].

Type material. Holotype: NHMW 1851 View Materials /0002/0035a, SL: 40.0 mm, MD: 25.8 mm, Grund ( Austria), illustrated in Hörnes (1853: pl. 24, figs 9a–b), Figs 20E View FIGURE 20 1 –E View FIGURE 1 3 View FIGURE 3 .

Illustrated material. NHMW 1851/0002/0035b, SL: 36.7 mm, MD: 22.5 mm, Grund ( Austria), holotype of Murex craticulatus erpis De Gregorio, 1885a and Murex craticulatus perisus De Gregorio, 1885a, illustrated in Hörnes (1853: pl. 24, figs 11a–b), Figs 20D View FIGURE 20 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 1851/0026/0026a, SL: 37.3 mm, MD: 24.6 mm, Grund ( Austria), Figs 20B View FIGURE 20 1 –B View FIGURE 1 3 View FIGURE 3 . NHMW 1851/0026/0026b, SL: 35.3 mm, MD: 24.1 mm, Grund ( Austria), Figs 20C View FIGURE 20 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 1859/0045/0121, SL: 37.8 mm, MD: 23.0 mm, Lectotype (designated herein) of Murex ( Occenebra [sic]) boeckhi Hoernes & Auinger, 1885 , Niederleis ( Austria), illustrated in Hoernes & Auinger (1885: pl. 27, fig. 3), Figs 20A View FIGURE 20 1 –A View FIGURE 1 2 View FIGURE 2 .

Additional material. 14 spec., NHMW 1851 View Materials /0002/0035, Grund ( Austria) .

Revised description. Large, solid, broad fusiform shell; apical angle ~60–65°. Protoconch unknown.Teleoconch of up to five whorls. Suture linear or shallowly undulating. Early teleoconch whorls convex with faint shoulder placed below mid-whorl. Axial sculpture of about ten narrow ribs, separated by slightly wider interspaces, overrun by weak adis, IP and abis and more prominent P1, P2, P3 with delicate s1, s2, forming cancellate pattern on last two spire whorls. Later teleoconch whorls with broad, weakly convex to weakly concave subsutural ramp, marked shoulder angulation placed short distance above abapical suture. Axial ribs prominent, forming blunt tubercles at shoulder. Last whorl attaining ~80% of total height, slightly inflated with broad subsutural ramp, angled periphery, weakly convex below, rapidly contracting at base. About nine broad ribs, fading over subsutural ramp. Subsutural ramp moderately prominent adis, IP and abis with single tertiary intercalated, adjoined by thread at each side; P1 weak, P2–P5 prominent, P6,ADP, MP weaker, s2–s5 forming weaker but prominent recessed cords. Secondary cords adjoined by one tertiary thread on each side. Secondary cords weakly scabrose; growth lines forming cancellate pattern in interspaces. Fasciole prominent, swollen, delimiting broad pseudumbilicus. Aperture wide, ovate; outer lip thickened by terminal varix, with crenulate edge, bearing the denticles ID trifid, D1 to D6 often bifid. Denticles close behind peristome. Anal canal moderately incised. Siphonal canal moderately short, narrow or fused, slightly deflected to the left, weakly dorsally recurved. Columellar callus forming broad rim, sharply delimited from base, erect at transition to siphonal canal.

Paratethyan synonyms. Four names have been established for this species. De Gregorio (1885a) introduced Murex craticulatus ergnapus De Gregorio, 1885a based on the specimen illustrated in Hörnes (1853: pl. 24, fig. 9) and Murex craticulatus perisus De Gregorio, 1885a for the specimen illustrated by Hörnes (1853: pl. 24, fig. 11). Murex craticulatus erpis De Gregorio, 1885a is an objective synonym of Murex craticulatus perisus. As first revisers we chose Murex ergnapus De Gregorio, 1885a as the name for this species. Murex ( Occenebra [sic]) boeckhi Hoernes & Auinger, 1885 is a subjective junior synonym of Ocenebra ergnapus (De Gregorio, 1885a) .

Discussion. Ocenebra ergnapus (De Gregorio, 1885a) is characterized by its biconical last whorl with marked shoulder angulation. This species was discussed by Hörnes (1853) and De Gregorio (1885a) in context with Murex craticulatus of Brocchi, 1814, described from the Pliocene of Italy [= Gracilipurpura craticuloides ( Vokes, 1964) ]. This would imply a placement of the Paratethyan species in Gracilipurpura Jousseaume, 1880 [type species Gracilipurpura craticulata ( Bucquoy & Dautzenberg, 1882) ; present-day, Mediterranean Sea]. Gracilipurpura craticulata , however, is slender fusiform with high spire and long, sometimes fused siphonal canal, shallow subsutural ramp and comparatively delicate, scabrose spiral cords.

Occurrences in the Eastern Paratethys described Iljina (1993) from the Karaganian and Konkian (Middle Miocene) of Kazakhstan differ in their shallower subsutural ramp, convex periphery and prominent P5 and P6. We have not examined the Eastern Paratethyan material and we are not certain about the placement of those specimens.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Grund ( Austria) (Hoernes & Auinger 1885); Vienna Basin: Niederleis ( Austria) (Hoernes & Auinger 1885); Pannonian Basin: Bánd, Várpalota ( Hungary) ( Kovács & Vicián 2024; Strausz 1966); Krka Basin: Vajndol ( Slovenia) ( Mikuž 2009).

NHMW	Naturhistorisches Museum, Wien

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Muricidae
Genus	Ocenebra