Ocenebra holubicensis ( Friedberg, 1912 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 52-53

Ocenebra holubicensis ( Friedberg, 1912)

Figs 13 , 22A–C

* Murex holubicensis Friedb. — Friedberg 1912: 172, pl. 11, fig. 4.

Murex holubicensis Friedb. var. subacarinata Friedb. — Friedberg 1912: 173, pl. 11, fig. 5.

Murex holubicensis Friedb. — Friedberg 1938: 587.

Murex holubicensis Friedb. — Friedberg 1938: 115.

[ Murex holubicensis ] var. subacarinata Friedb. — Friedberg 1938: 116.

Muricopsis holubicensis Friedb. — Korobkov 1955: plate captions, pl. 70, figs 10a–b.

Muricopsis holubicensis ( Friedberg, 1912) — Zelinskaya et al. 1968: 187, pl. 44, figs 15–16.

Murex holubicensis Friedberg — Bałuk 1970: 118, pl. 12, figs 3a–2b.

[ Muricopsis ] holubicensis Friedberg, 1912 — Vokes 1971: 58.

Muricopsis holubicensis ( Friedberg, 1912) — Stojaspal 1978: 334.

Type material. Holotype: Nr. 3608, State Natural History Museum, National Academy of Sciences of Ukraine, L’viv), SL: 23.2 mm, MD: 11.5 mm, Hołubica ( Holubytsia ) ( Ukraine), illustrated in Friedberg (1912: pl. 11, fig. 4), Fig. 22A .

Illustrated material Lectotype of Murex holubicensis subacarinata Friedberg, 1912 (designated herein): Nr.: 3607-1c, State Natural History Museum, National Academy of Sciences of Ukraine, L’viv), SL: 13.1 mm, MD: 10.0 mm, Hołubica (Holubytsia) ( Ukraine), illustrated in Friedberg (1912: pl. 11, fig. 5), Fig. 22B . Paralectotype: of Murex holubicensis subacarinata Friedberg, 1912: Nr.: 3607-2b, State Natural History Museum, National Academy of Sciences of Ukraine, L’viv), SL: 18.0 mm, MD: 8.9 mm, Hołubica (Holubytsia) ( Ukraine), Fig. 22C .

Revised description. Medium-sized, slender fusiform shell with high spire; apical angle ~50°. Protoconch unknown. Teleoconch of up to six whorls. Suture moderately incised, shallowly undulating. Early teleoconch whorls with broad, weakly concave subsutural ramp, angled shoulder, subcylindrical periphery, constricted below. Axial sculpture of eight prominent, wide-spaced ribs. adis, IP, abis weak, P1 and P2 prominent, slightly swollen over ribs; P1 weakly tuberculate; s1, s2 delicate. Last whorl attaining ~70% of total height; with concave subsutural ramp, weakly angled shoulder, weakly convex periphery, moderately constricted below, with conical base. Eight prominent axial ribs fading over subsutural ramp, persisting onto base. P1–P6 forming prominent, moderately narrow, rounded cords separated by wider interspaces; s1–s6 delicate; ADP, MP more prominent, close-set. Fasciole moderately swollen, relatively narrow, delimiting narrow, shallow pseudoumbilicus. Aperture narrowly ovate. Outer lip thickened with prominent D1–D6. Anal canal moderately incised, poorly delimited. Siphonal canal moderately long, open, moderately narrow, slightly bent to the left. Columella straight, twisted at siphonal canal. Columellar callus forming broad rim, erect at siphonal canal, adherent in parietal area.

Paratethyan synonyms. Murex holubicensis subacarinata Friedberg, 1912 was introduced as subspecies based on its rounded shoulder.

Discussion. This species was listed as Muricopsis by Korobkov (1955) and Vokes (1971). This placement is rejected based on the apertural features. Muricopsis species typically have a prominent ID, an atrophied D1 and a large, knob-like D2 followed by weaker D3–D5. In addition, typical Muricopsis has one or two columellar folds, which are absent in Ocenebra holubicensis ( Friedberg, 1912) .

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Voronyaky Hills: Hołubica (Holubytsia), Podhorce (Pidhirtsi) ( Ukraine). Polish-Ukrainian Fore-Carpathian Basin: Bogucice, Hołdy, Faszczówka, Stary Poczajów, Tarnoruda, Zaleśce (Salisze), Żukowce (Zhukivtsi), Zborów (Zboriv) ( Ukraine) ( Friedberg 1912; 1938).

Figures

FIGURE 13. Cord diagrams and apertural features of Ocenebrinae [Ceratostoma? steiningeri sp. nov., Gracilipurpura polonica (Bałuk, 1995), Heteropurpura catosus (De Gregorio, 1885a), Kestocenebra subaustriaca (Stojaspal, 1978), Kestocenebra vermeiji sp. nov., Kestocenebra? globosa (Stojaspal, 1978), Ocenebra ergnapus (De Gregorio, 1885a), Ocenebra haidingeri (Hörnes, 1853), Ocenebra holubicensis (Friedberg, 1912), Ocenebra littoralis nov. nom., Ocenebra mirmigus (De Gregorio, 1885a), Ocenebra scorpio sp. nov., Ocenebra valdemari (Bałuk, 2006), Ocenebra vindobonensis (Hörnes, 1853), Ocenebra zboroviensis (Friedberg, 1912)].

FIGURE 22. Ocenebra holubicensis (Friedberg, 1912). A. holotype: Nr. 3608, Hołubica (Holubytsia) (Ukraine). B. Nr.: 3607- 1c, Hołubica (Holubytsia) (Ukraine). C. Nr.: 3607-2b, Hołubica (Holubytsia) (Ukraine).