Ocenebra mirmigus (De Gregorio, 1885a)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 55-56

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FFD0-806D-FF50-ACF9F963B8F8
treatment provided by	Plazi
scientific name	Ocenebra mirmigus (De Gregorio, 1885a)
status

Treatment

Figs 13 View FIGURE 13 , 24A–D View FIGURE 24

[ Murex View in CoL ] Lassaignei Bast. View in CoL — Hörnes 1848: 18 [non Ocenebra lassaignei (de Basterot, 1825) View in CoL ].

Murex Lassaignei Bast. View in CoL —Hörnes 1853: 232, pl. 24, fig. 8 [non Ocenebra lassaignei (de Basterot, 1825) View in CoL ].

* [ Murex Lassaignei Bast. View in CoL ] Var. mirmigus De Greg. —De Gregorio 1885a: 249 [pro Murex lassaignei sensu Hörnes 1853: 232 View in CoL , pl. 24, fig. 8].

Murex (Phyllonotus) Lassaignei Bast. View in CoL —Hoernes & Auinger 1885: 213 [non Ocenebra lassaignei (de Basterot, 1825) View in CoL ].

Murex (Phyllonotus) Vindobonensis M. Hoern. View in CoL —Hoernes & Auinger 1885: 213, pl. 25, fig. 2 [non Ocenebra vindobonensis (Hörnes, 1853) View in CoL ].

P [urpura]. (T [ritonalia].) lassaignei (Bast.) — Sieber 1958: 147.

Tritonalia (Tritonalia) lassaignei Basterot 1825 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 149, pl. 30, fig. 8 [non Ocenebra lassaignei (de Basterot, 1825) View in CoL ].

Tritonalia vindobonensis (Hörnes) var.— Kókay 1966: 57, pl. 8, fig. 6 [non Ocenebra vindobonensis (Hörnes, 1853) View in CoL ].

[ Ocenebra View in CoL ] mirmigus De Gregorio 1885 — Vokes 1971: 71.

Ocenebra (Ocenebra) lassaignei ( Basterot, 1825) View in CoL — Stojaspal 1978: 338 [non Ocenebra lassaignei (de Basterot, 1825) View in CoL ].

Purpura (Tritonalia) kojumdgievae View in CoL nom. n. — Bałuk 1995: 228, pl. 30, fig. 8.

Ocinebrina kojumdgievae ( Bałuk, 1995) View in CoL — Landau et al. 2013: 151, pl. 22, fig. 6.

Ocinebrina kojumdgievae ( Bałuk, 1995) View in CoL — Goret & Pons 2013: 60, pl. 3, fig. 3.

Ocinebrina kojumdgievae ( Bałuk, 1995) View in CoL — Vicián et al. 2017: 269, pl. 2, figs 11–12.

Ocinebrina kojumdgievae ( Bałuk, 1995) View in CoL — Kovács 2019: 127, figs 48–49.

Ocinebrina kojumdgievae ( Bałuk, 1995) View in CoL — Kovács 2020: 456, pl. 2, figs 27–30.

? Ocinebrina cf. kojumdgievae ( Bałuk, 1995) View in CoL — Kovács et al. 2023: 26, fig. 63.

Type material. Holotype: NHMW 1849 View Materials /0005/0022, SL: 30.7 mm, MD: 20.3 mm, Bad Vöslau ( Austria), illustrated in Hörnes (1853: pl. 24, fig. 8), Figs 24A View FIGURE 24 1 –A View FIGURE 1 3 View FIGURE 3 .

Illustrated material. NHMW 1865/0001/0190a, SL: 36.6 mm, MD: 21.2 mm, Lăpugiu de Sus ( Romania), Figs 24B View FIGURE 24 1 –B View FIGURE 1 3 View FIGURE 3 . NHMW 1865/0001/0190b, SL: 39.9 mm, MD: 23.5 mm, Lăpugiu de Sus ( Romania), Figs 24C View FIGURE 24 1 – C View FIGURE 1 2 View FIGURE 2 . NHMW 1865/0001/0191, SL: 29.9 mm, MD: 19.2 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1885: pl. 25, fig. 2), Figs 24D View FIGURE 24 1 –D View FIGURE 1 2 View FIGURE 2 .

Additional material. 3 spec., NHMW 1846 View Materials /0037/0233, Enzesfeld ( Austria) ; 12 spec., NHMW 1974 View Materials /0025/0029, Lăpugiu de Sus ( Romania) ; 1 spec., NHMW 1867 View Materials /0019/0091, Coşteiu de Sus ( Romania) .

Revised description. Medium-sized, very solid, stocky shell with cyrtoconoid early spire; apical angle ~90°, decreasing to ~60° on later spire whorls. Protoconch unknown. Teleoconch of up to five whorls. Early teleoconch whorls low, with very broad, shallow subsutural ramp; shoulder placed just above abapical suture. About ten tubercles along shoulder. High, thin subsutural collar developing on 3 rd to 4 th teleoconch whorl; subsutural ramp becoming steep and strongly concave with numerous flattened spiral cords. Last whorl attaining ~85% of total height, with high subsutural collar, concave subsutural ramp, angled shoulder, weakly convex below, strongly constricted at base. Axial sculpture of four abapically weakening varices with one only slightly weaker intervarical rib; terminal varix flaring, foliaceous. Spiral sculpture: adis, IP, abis, delicate adjoined by subequal tertiary cords; P1–P5 forming prominent, rounded, subequal cords, occasionally terminating in short, pointed tubercles on varices (see Kovács 2020: pl. 2, figs 27–30); P3 sometimes weak; P1 forming trigonal tubercles at shoulder varices; s1–s6 delicate adjoined by tertiary thread on either side; P6, ADP and MP weak. Delicately scabrose growth lines in spiral interspaces. Fasciole moderately swollen, delimiting very broad, shallow pseudoumbilicus. Aperture narrowly ovate; outer lip strongly thickened with very prominent ID, D1–D5, rounded denticles loosely arranged in pairs close behind peristome. Anal canal distinctly incised, moderately narrow; siphonal canal moderately long, fused, straight to slightly bent to the right. Columella straight, smooth, weakly twisted at transition in siphonal canal. Columellar callus forming broad adherent rim, sharply delimited from base.

Paratethyan synonyms. Bałuk (1995) introduced Purpura (Tritonalia) kojumdgievae as new name for the Paratethyan specimens, which had been erroneously treated as Ocinebrina lassaignei (de Basterot, 1825) in the literature. However, Bałuk (1995) overlooked that De Gregorio (1885a) had already introduced Murex lassaignei mirmigus for this species. Therefore, Purpura (Tritonalia) kojumdgievae is an objective junior synonym of Ocenebra mirmigus (De Gregorio, 1885a) .

Discussion. This species is characterized by the allometric growth of its spire, which develops a high subsutural collar and a prominent very broad terminal varix, which is strongly flaring in its adapical half, causing a somewhat subquadratic outline in apertural view. Ocenebra scorpio sp. nov. and Ocenebra vindobonensis (Hörnes, 1853) differ in their prominent spines on the varices. The mode of formation of primary cords, with reduced P2, is comparable to Ocenebra scorpio sp. nov. Specimens of Ocinebrina mirmigus have been incorrectly assigned to Ocinebrina lassaignei (de Basterot, 1825) by several authors since Hörnes (1853). However, they differ quite distinctly in the shorter spire, more convex whorls, weaker shoulder and more prominent axial ribs in O. lassaignei , which lacks a flaring terminal varix (see Lozouet et al. 2001: pl. 23, fig. 8; Goret & Pons 2013: pl. 3, fig. 2). This species group is present from the Lower Miocene to the Pliocene, last represented in the Mediterranean by O. ariesiensis ( Fontannes 1879) . The shells of species within this group are very similar and need to be revised.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys.? Ottnangian (Early Miocene): Early Miocene North Hungarian Basin: Parádfürdö ( Hungary) ( Kovács et al. 2023). Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1995); Vienna Basin: Bad Vöslau, Enzesfeld ( Austria) (Hoernes & Auinger 1885); Bakony Mountains: Bánd ( Hungary) ( Vicián et al. 2017); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus ( Romania) ( Kovács 2019); Dacian Basin: Staropatica, Târnene, Radomirci ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Seyithasan ( Turkey) ( Landau et al. 2013).

NHMW	Naturhistorisches Museum, Wien

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Muricidae
Genus	Ocenebra