Exquisitor graniferus ( Michelotti, 1841 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 33-35

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FFC6-8078-FF50-AE32F92ABEF9
treatment provided by	Plazi
scientific name	Exquisitor graniferus ( Michelotti, 1841 )
status

Treatment

Exquisitor graniferus ( Michelotti, 1841) View in CoL

Figs 1 View FIGURE 1 , 12A–C View FIGURE 12

* Murex graniferus View in CoL mihi— Michelotti 1841: 11, pl. 5, fig. 6.

Murex graniferus Micht. View in CoL —Hörnes 1853: 254, pl. 26, figs 1a–c. Murex graniferus Michtti View in CoL — Bellardi 1873: 65, pl. 5, figs 1a–b.

[ Murex torularius Lamarck View in CoL ] F. a pirlus —De Gregorio 1885a: 230 [pro Murex graniferus sensu Hörnes 1853 View in CoL : pl. 26, fig.1]. Murex (Pteronotus) graniferus Michti. View in CoL —Hoernes & Auinger 1885: 204.

Murex View in CoL ( Tubicauda View in CoL ) subgranifer Cossmann & Peyrot 1924: 99, no.725, pl. 17, fig. 34, pl. 18, fig. 2.

M [urex]. (T [ubicauda].) graniferus Micht. View in CoL — Sieber 1958: 145.

Murex View in CoL ( Tubicauda View in CoL ) subgranifer Cossmann & Peyrot— Erünal-Erentöz 1958: 54, pl. 8, figs 3–4.

[ Subpterynotus View in CoL ] graniferus Michelotti, 1841 View in CoL — Vokes 1971: 55.

Murexiella (Subpterynotus) graniferus ( Michelotti, 1841) — Stojaspal 1978: 334.

Subpterynotus graniferus (Michelotti) — Landau et al. 2013: 162, pl. 24, figs 6–7.

Subpterynotus graniferus (Michelotti) — Vicián et al. 2017: 270, pl. 2, figs 17–18.

Subpterynotus graniferus ( Michelotti, 1841) — Kovács et al. 2018: 120, figs 5M–N.

Exquisitor graniferus Merle et al. 2022: 161 View in CoL , 478, pl. 155, figs 1–6.

Type material. The spire fragment illustrated by Michelotti (1841, pl. 5, fig. 6) from the Tortonian of S. Agata ( Italy) might be stored in the Museo Regionale di Scienze Naturali Torino ( Italy) but was not listed by Ferrero Mortara et al. (1981: 32).

Illustrated material. NHMW 1876/0011/0030, SL: 56.5 mm, MD: 32.1 mm, Lăpugiu de Sus ( Romania), Figs 12A View FIGURE 12 1 –A View FIGURE 1 3 View FIGURE 3 . NHMW 1846/0037/0061, SL: 67.4 mm, MD: 37.9 mm, Gainfarn ( Austria), Figs 12B View FIGURE 12 1 –B View FIGURE 1 3 View FIGURE 3 . NHMW 1847/0037/0063, SL: 46.9 mm, MD: 31.6 mm, Weinsteig ( Austria), illustrated in Hörnes (1853: pl. 26, fig. 1), note that Hörnes (1853) gave Forchtenau (Badenian) as locality, but Weinsteig (Karpatian) is indicated on the labels, Figs 12C View FIGURE 12 1 –C View FIGURE 1 2 View FIGURE 2 .

Additional material. 1 spec., NHMW 1846 View Materials /0037/0211, Enzesfeld ( Austria) ; 1 spec., NHMW 1858 View Materials /0033/0031, Lăpugiu de Sus ( Romania) ; 2 spec., NHMW 1867 View Materials /0029/0084, Coşteiu de Sus ( Romania) ; 1 spec., NHMW A259 View Materials , Nemeșești ( Romania) .

Revised description. Medium-sized, solid, trivaricate, club-shaped shell with long siphonal canal; apical angle ~50°. Protoconch unknown. Early teleoconch whorls weakly shouldered with convex subsutural ramp. Suture moderately impressed, shallowly undulating. Axial sculpture of three prominent, raised, trigonal varices and three weaker intervarical ribs. Spiral sculpture on antepenultimate and penultimate whorls of moderately prominent adis, IP and abis with single tertiary thread intercalated in interspaces. Weak P1–P3 with only slightly weaker s1–s3. Spiral cords and interspaces slightly scabrose in well preserved specimens. P1 forming short spines on shoulder varices on last two spire whorls. Last whorl attaining 75–80% of total height, with broad, convex, shallow, subsutural ramp delimited by angular shoulder, moderately convex below, strongly constricted at base. Subsutural ramp with weak adis, IP and abis with tertiary cords and two additional threads intercalated in interspaces. P1 most prominent forming short spines on shoulder varices. P2 and P3 weaker, P4 almost as strong as P1 delimiting base, P5 and P6 weak, forming short spines on terminal varix. s1–s6 narrow, with up to three tertiary spiral threads intercalated in interspaces. ADP and MP of variable strength, often forming terminal spines; ABP and EAB1 occasionnaly present without forming terminal spines. Aperture narrowly ovate. Outer lip smooth edged and smooth within. Anal canal subobsolete. Siphonal canal long, moderately narrow, straight, weakly deflected to the left, shallowly notched. Columella strongly excavated in adapical half, smooth, markedly angulated at transition to siphonal canal. Columellar callus forming narrow thickened, adherent rim, sharply delimited from base.

Paratethyan synonyms. Murex torularius pirlus De Gregorio, 1885a was introduced by De Gregorio (1885a) for the specimen illustrated in Hörnes (1853: pl. 26, fig.1). We treat this taxon as subjective junior synonym of Subpterynotus graniferus ( Michelotti, 1841) .

Discussion. This species was placed by Vokes (1971) in Subpterynotus Olsson & Harbison, 1953 [type species Murex (Pteronotus) textilis Gabb, 1873 ; Miocene, Dominican Republic]. However, Merle et al. (2022) showed it to be unrelated to that Caribbean genus. In addition, they showed that the sculptural pattern of E. graniferus and E. exquisitus shares similarities with that found in Siratus belonging to the Muricinae , but displays differences that justify a generic distinction. Thus, they erected the genus Exquisitor with the West African Murex exquisitus G.B. Sowerby III, 1904 as type species.

Paleoenvironment. Shallow marine, inner neritic, along mudflats (Weinsteig: Harzhauser et al. 2002), in seagrass beds (Gainfarn: Zuschin et al. 2007) and in the vicinity of corals (Letkés: Kovács & Vicián 2014).

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Weinsteig ( Austria) (Hoernes & Auinger 1885). Badenian (Middle Miocene): Vienna Basin: Gainfarn, Enzesfeld ( Austria), Boršov (= Porstendorf) ( Czech Republic) (Hoernes & Auinger 1885); Pannonian Basin: Letkés ( Hungary) ( Kovács et al. 2018); Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus, Nemeşeşti ( Romania) (Hoernes & Auinger 1885).

Proto-Mediterranean Sea. Burdigalian (Early Miocene): Colli Torinesi: Baldissero ( Italy) ( Bellardi 1873); Serravallian (Middle Miocene): Karaman Basin: Lale, Akpınar, Seyithasan ( Turkey) ( Landau et al. 2013); Crete, Heraklio Basin: Tilissos ( Greece) ( Merle et al. 2022); Tortonian (Late Miocene): Po Basin: S. Agata ( Italy) ( Bellardi 1873).

Northeastern Atlantic. Burdigalian (Early Miocene): Aquitaine Basin: Léognan, Saucats ( France) ( Cossmann & Peyrot 1924).

NHMW	Naturhistorisches Museum, Wien

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Muricidae
Genus	Exquisitor