Ocenebra confluens ( Eichwald, 1830 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 65-66

Ocenebra confluens ( Eichwald, 1830)

Figs 13 , 28C–E

* M [urex]. confluens m.— Eichwald 1830: 224.

Murex confluens m.— Eichwald 1851: 101, pl. 8, fig. 11.

Mur [ex]. confluens — Eichwald 1852: plate captions, pl. 8, fig. 11.

Mur [ex]. confluens m.— Eichwald 1853: 192.

M [urex]. Galicianus sp. nov. — Hilber 1881: 184 [nomen nudum].

Murex Galicianus Hilber, nova forma — Hilber 1882: 5 (pars), pl. 1, figs 7a–b [non figs 8a–c = Orania agapus (De Gregorio, 1885a) ].

Murex ( Chicoreus ) Galicianus Hilb.—Hoernes & Auinger 1885: 209 (pars), pl. 29, figs 12a–c [non figs 11a–c = Orania agapus (De Gregorio, 1885a) ].

Murex confluens Eichw. — Friedberg 1912: 166, pl. 10, figs 6–7.

Murex confluens Eichw. var. convexus Friedb. — Friedberg 1912: 167, pl. 10, fig. 8.

Murex confluens Eichw. — Friedberg 1938: 115.

Tritonalia (Hadriania) aff. mioincrassata — Csepreghy-Meznerics 1950: 48, pl. 2, fig. 19.

Murex confluens Eichwald — Zelinskaya et al. 1968: 185, pl. 44, figs 5–6.

Murex confluens var. convexus Friedberg 1912 — Zelinskaya et al. 1968: 186.

Murex confluens Eichwald — Bałuk 1970: 118, pl. 12, figs 2a–2b.

[ Ocinebrina ] confluens Eichwald, 1830 — Vokes 1971: 34.

[ Ocinebrina ] convexus ( Friedberg, 1912 — Vokes 1971: 35.

? Murex ( Chicoreus ) galicianus Hilber an sp. dist.— Csepreghy-Meznerics 1969: 81, pl. 4, fig. 26.

Tritonalia confluens Eichwald — Csepreghy-Meznerics 1972: 27, pl. 10, fig. 6.

Ocenebra (Ocinebrina) confluens ( Eichwald, 1830) — Stojaspal 1978: 335.

Ocenebra (Ocinebrina) convexa ( Friedberg, 1912) — Stojaspal 1978: 335.

Ocenebra (Ocinebrina) galiciana ( Hilber, 1882) — Stojaspal 1978: 335.

Murex confluens Eichwald — Ionesi & Nicorici 1994: pl. 3, fig. 7.

Purpura sp. — Bałuk 1995: 228

Purpura (Tritonalia) confluens ( Eichwald, 1853) — Bałuk 2006: 209, pl. 13, figs 1–3.

Ocinebrina confluens ( Eichwald, 1853) — Kovács 2020: 456, pl. 2, figs 24–25.

non Murex ( Chicoreus ) galicianus Hilber an sp. dist.— Csepreghy-Meznerics 1972: 27, pl. 9, fig. 26 [= Ocinebrinopsis credneri or Ocinebrinopsis dominicii group].

non Ocinebrina confluens ( Eichwald, 1853) — Švagrovský 1982: 387, pl. 1, fig. 6 [= Ocinebrinopsis credneri (Hoernes & Auinger, 1885) ].

non Murex confluens Eichw. (? subsp. nov.)—Nevesskaya et al. 1986: 172, pl. 8, figs 2–3 [= another species of the Ocenebra schoenni group].

non Murex (Tritonalia) confluens (Eichwald) — Jovanović 2010: 84, text-fig. 1.2 [=? Murexsul sp. ].

Type material. Lectotype designated by Švagrovský (1982: 387) illustrated in Eichwald (1852: pl. 8, fig. 11); SL: 20.6 mm, MD: 10.1 mm (according to illustrations in Eichwald (1851, 1852), Biłka or Żukowce (Zhukivtsi) ( Ukraine). The specimen could not be reliably identified in the St. Petersburg State University, Paleontological Museum ( Russia) ( Fig. 28D ).

Holotype of Murex confluens convexus Friedberg, 1912: Nr. 3610, State Natural History Museum, National Academy of Sciences of Ukraine, L’viv), SL: 19.2 mm, MD: 9.2 mm, Zborów (Zboriv) ( Ukraine), illustrated in Friedberg (1912: pl. 10, fig. 8), Fig. 28E ).

The three syntypes of Murex galicianus Hilber, 1882 were stored in the Geological Survey in Vienna but were destroyed during WWII .

Illustrated material. Collection of Zoltán Kovács, SL: 19 mm, MD: 10 mm, Hidas ( Hungary), illustrated in Kovács 2020: 456, pl. 2, figs 24–25; Figs 28C 1 –C 2 .

Revised description. Small, broadly fusiform shell with high, gradate spire; apical angle ~55°. Protoconch unknown. Teleoconch of up to six whorls; suture deeply incised, shallowly undulating. Early teleoconch whorls with shallow, weakly concave subsutural ramp and shoulder placed slightly above mid-whorl. Axial sculpture of about eight prominent ribs overrun by delicate adis, IP, abis, slightly more prominent P1 along shoulder, subequal P2 and delicate s1 and s2; P3 appears at abapical suture on third teleoconch whorl. Last whorl attaining ~70% of total height; slightly allometrically widening with broad, shallow subsutural ramp, marked shoulder, weakly convex periphery, moderately constricted below. Axial sculpture of three broad varices and up to two weaker intervarical ribs. Spiral sculpture of prominent P1–P5 on convex part of whorl; P6 atrophied on the siphonal canal; on subsutural ramp adis, IP, adis weak, ADP, MP, ABP weak; s1–s6, ads delicate, recessed, adjoined by tertiary thread on either side. Fasciole swollen, twisted, delimiting deep, moderately narrow pseudoumbilicus. Aperture narrow, elongate ovate; outer lip strongly thickened, nearly straight-sided, with very prominent D1–D5. Anal canal weakly incised; siphonal canal moderately long, fused in adult specimens, slightly bent to the left. Columella straight, smooth. Columellar callus forming broad, sharply delimited rim, erect at siphonal canal, adherent in parietal area.

Paratethyan synonyms. Hilber (1882) illustrated two syntypes of Murex galicianus Hilber, 1882 from Hołubica ( Ukraine). The specimen illustrated on pl. 1, fig. 7 is indistinguishable from Ocenebra confluens ( Eichwald, 1830) , whereas fig. 8 represents Orania agapus . As first revisers we select the specimen illustrated by Hilber (1882: pl. 1, fig. 7) as lectotype of Murex galicianus, which makes this species a subjective synonym of Ocenebra confluens ( Eichwald, 1830) . The material described by Hilber (1882) was stored in the collection of the Geological Survey in Vienna (now Geosphere Austria) but was destroyed during World War II. Ocinebrina confluens convexa ( Friedberg, 1912) is based on a single poorly preserved specimen that differs from typical Ocenebra confluens ( Eichwald, 1830) by its convex periphery. Further material would be needed to evaluate the status of this taxon. Herein, we tentatively treat it a subjective junior synonym of O. confluens , with which it co-occurs.

Discussion. Ocenebra schoenni (Hörnes, 1853) has an identical shape and very similar apertural features like Ocenebra confluens . However, they differ mainly in the cancellate sculpture of Ocenebra schoenni .

Occurrences in the Eastern Paratethys described by Nevesskaya et al. (1986) from the Konkian (Middle Miocene) of Kazakhstan differ in their cancellate sculpture and are closer to Ocenebra schoenni (Hörnes, 1853) . The two specimens illustrated by Nevesskaya et al. (1986) lack apertural features and their definitive identification is impossible .

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1995); Nowy Sącz Basin: Niskowa ( Poland) ( Bałuk 1970); Polish-Ukrainian Fore-Carpathian Basin: Biłka, Hołdy, Jasinów, Olesko, Płuhów, Zborów (Zboriv), Żukowce (Zhukivtsi) ( Ukraine) ( Eichwald 1830; Friedberg 1912, 1938); Voronyaky Hills: Hołubica (Holubytsia), Podhorce (Pidhirtsi) ( Ukraine) ( Friedberg 1912); Pannonian Basin: Hidas ( Hungary) ( Kovács 2020); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972).

Figures

FIGURE 1. Cord diagrams and apertural features for Haustrinae [Haustrum? pyrulatum (Hoernes & Auinger, 1882)] and Muricinae [Bolinus gapus (De Gregorio, 1885a), Bolinus partschi (Hörnes, 1853), Chicoreus (Triplex) aquitanicus (Grateloup, 1833), Chicoreus (Triplex) hirmetzli (Kovács, 2018), Chicoreus (Triplex) notatus (Eichwald, 1830). Hexaplex (Trunculariopsis) trunculoides (Pusch, 1837), Hexaplex (Trunculariopsis) esplus (De Gregorio, 1885a), Hexaplex(Trunculariopsis) subasperrimus (d’Orbigny, 1852), Promurex spinicosta (Bronn, 1831), Exquisitor graniferus (Michelotti, 1841)].

FIGURE 2. A1–A2. Haustrum? pyrulatum (Hoernes & Auinger, 1882), holotype, NHMW 1859/0045/0590, Lăpugiu de Sus (Romania). B1–B2. Haustrum lacunosum (Bruguière, 1789), MNHN-IM-2000-904, lectotype, New Zealand, present-day, photo: Manuel Caballer, 2014 RECOLNAT project ANR-11-INBS-0004.

FIGURE 13. Cord diagrams and apertural features of Ocenebrinae [Ceratostoma? steiningeri sp. nov., Gracilipurpura polonica (Bałuk, 1995), Heteropurpura catosus (De Gregorio, 1885a), Kestocenebra subaustriaca (Stojaspal, 1978), Kestocenebra vermeiji sp. nov., Kestocenebra? globosa (Stojaspal, 1978), Ocenebra ergnapus (De Gregorio, 1885a), Ocenebra haidingeri (Hörnes, 1853), Ocenebra holubicensis (Friedberg, 1912), Ocenebra littoralis nov. nom., Ocenebra mirmigus (De Gregorio, 1885a), Ocenebra scorpio sp. nov., Ocenebra valdemari (Bałuk, 2006), Ocenebra vindobonensis (Hörnes, 1853), Ocenebra zboroviensis (Friedberg, 1912)].

FIGURE 28. A–B. Ocenebra zboroviensis (Friedberg, 1912). A1–A2. lectotype, Zborów (Zboriv) (Ukraine), illustrated in Friedberg (1912: pl. 10, figs 4a–b). B1–B2. paralectotype: Zborów (Zboriv) (Ukraine), illustrated in Friedberg (1912: pl. 10, figs 5a–b), C–E. Ocenebra confluens (Eichwald, 1830). C1–C2. collection of Zoltán Kovács, Hidas (Hungary). D. lectotype illustrated in Eichwald (1852: pl. 8, fig. 11), Biłka or Żukowce (Zhukivtsi) (Ukraine). E. Nr. 3610, Zborów (Zboriv) (Ukraine). F–H. Ocenebra schoenni (Hörnes, 1853). F1–F2. Paralectotype, NHMW 1852/0012/0009b, Maria Dreieichen (Austria). G1–G2. Lectotype, NHMW 1852/0012/0009a, Maria Dreieichen (Austria). H. NHMW 1852/0012/0009c, Maria Dreieichen (Austria).