Ocinebrinopsis gregaria, Harzhauser & Landau & Merle, 2025

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 71-73

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FFA0-8012-FF50-AFACFBE1B843
treatment provided by	Plazi
scientific name	Ocinebrinopsis gregaria
status	sp. nov.

Treatment

Ocinebrinopsis gregaria sp. nov.

Figs 29 View FIGURE 29 , 31A–F View FIGURE 31

Murex sublavatus Basterot View in CoL —Hörnes 1853: 236 (pars), pl. 24, fig. 16 [non Ocenebra sublavata (de Basterot, 1825) View in CoL ].

Murex View in CoL ( Occenebra [sic]) caelatus Grat.—Hoernes & Auinger 1885: 217 (pars), pl. 26, figs 7–12 [non Ocinebrinopsis coelata ( Dujardin, 1837) ].

Tritonalia (Tritonalia) credneri (H.-Au.)— Csepreghy-Meznerics 1954: 37, pl. 4, figs 6, 11, 19 [non Ocinebrinopsis credneri (Hoernes & Auinger, 1885) ].

O [cinebrina]. caelata Grat. — Sieber 1958: 146 [non Ocinebrinopsis coelata ( Dujardin, 1837) ].

Ocinebrina caelata (Grat.) — Kókay 1966: 57, pl. 8, figs 3–4 [non Ocinebrinopsis coelata ( Dujardin, 1837) ].

? Ocinebrina sublavata sublavata ( Basterot, 1825) View in CoL —Hinculov in Iliescu et al. 1968: 140, pl. 34, figs 7a–b [non Ocenebra sublavata (de Basterot, 1825) View in CoL ].

Hadriania excoelata ( Cossmann & Peyrot, 1924) — Bałuk 1995: 224, pl. 28, figs 2–3, pl. 29, fig. 1 [non Ocinebrina excoelata ( Cossmann & Peyrot, 1924) ].

Type material. Holotype. NHMW 1949 View Materials /0005/0021a, SL: 22.0 mm, MD: 13.1 mm, Enzesfeld ( Austria), illustrated in Hoernes & Auinger (1885: pl. 26, fig. 11), Figs 31B View FIGURE 31 1 –B View FIGURE 1 3 View FIGURE 3 . Paratypes: NHMW 2024 View Materials /0046/0001, SL: 21.2 mm, MD: 12.8 mm, Enzesfeld , Gainfarn or Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1885: pl. 26, fig. 9), Figs 31A View FIGURE 31 1 –A View FIGURE 1 3 View FIGURE 3 . NHMW 1863 View Materials /0015/1122a, SL: 21.6 mm, MD: 12.6 mm, Gainfarn ( Austria), Figs 31C View FIGURE 31 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 1863 View Materials /0015/1122b, SL: 19.8 mm, MD: 13.4 mm, Gainfarn ( Austria), Figs 31D View FIGURE 31 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 1949 View Materials /0005/0021b, SL: 22.3 mm, MD: 12.4 mm, Enzesfeld , Gainfarn or Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1885: pl. 26, fig. 10), Figs 31E View FIGURE 31 1 –E View FIGURE 1 3 View FIGURE 3 . NHMW 1949 View Materials /0005/0021b, SL: 20.7 mm, MD: 10.7 mm, Enzesfeld , Gainfarn or Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1885: pl. 26, fig. 7), Figs 31F View FIGURE 31 1 –F View FIGURE 1 2 View FIGURE 2 .

Additional material. 40 spec., NHMW 1855 View Materials /0045/0436, Gainfarn ( Austria) ; 65 spec., 1846/0037/0226, Enzesfeld and Gainfarn ( Austria) .

Type locality. Enzesfeld ( Austria), Vienna Basin .

Type stratum. Baden Formation.

Age. Middle Miocene, middle Badenian (Langhian).

Etymology. gregarius, Latin for living in flocks, in a group, being common; referring to the abundant occurrence.

Diagnosis. Medium-sized, stout fusiform shell with inflated last whorl, prominent fasciole delimiting deep pseudoumbilicus, fused siphonal canal and lirae in outer lip.

Description. Medium-sized, stout fusiform to globose shell with conical spire; apical angle ~60°. Protoconch unknown. Teleoconch of up to five whorls. Suture weakly incised, shallowly undulating. Early teleoconch whorls convex with convex to weakly concave subsutural ramp and rounded shoulder. Axial sculpture of about nine broad, prominent ribs, slightly weakening over subsutural ramp. Spiral sculpture on antepenultimate and penultimate whorls of weakly scabrose adis, IP and abis; P1, s1, P2, s2 of subequal strength. Last whorl inflated, attaining ~75–80 of total height; relatively shallow, weakly concave subsutural ramp, weakly angled to rounded shoulder. Convex periphery, strongly constricted below. About ten broad, weak axial ribs. Spiral sculpture with weakly scabrose subsutural threads, prominent adis, IP and abis, P1–P6 of almost equal strength; P6 on convex part of base; adis, P1, P5, P6 slightly more prominent; s1–s4, s6 slightly weaker, slightly recessed, s5 prominent; ADP, MP and ABP present; one slightly scabrose tertiary thread may appear in interspaces. Fasciole strongly swollen, twisted, delimiting broad, shallow pseudoumbilicus. Aperture ovate; outer lip strongly thickened, with weakly crenulate edge; ID, D1–D6 lirate, starting close behind peristome; denticles occasionally bifid. Anal canal weakly incised; siphonal canal moderately long, narrow, fused, straight, slightly dorsally recurved tip. Columella straight, smooth, slightly twisted at siphonal canal. Columellar callus forming broad rim, sharply delimited, weakly erect at siphonal canal, adherent in parietal area.

Discussion. Ocinebrinopsis gregaria sp. nov. is a common species in shallow water deposits at Gainfarn and Enzesfeld ( Austria). It displays some variability in profile, ranging from globose specimens to slightly more slender ones with a relatively high spire. Since Hoernes & Auinger (1885), it has been confused with Ocinebrinopsis coelata ( Dujardin, 1837) from the Middle Miocene of the Loire Basin. This misidentification is difficult to understand, as O. coelata is very slender, has a much narrower aperture and has rounded denticles within the outer lip rather than short lirae (see Landau et al. 2019, pl. 25, figs 1–3 for O. coelata ).

Bałuk (1995) identified this Paratethyan species with Ocinebrina excoelata ( Cossmann & Peyrot, 1924) from the Burdigalian of Léognan ( France). Lozouet et al. (2001) proposed a very wide species concept for Ocinebrina excoelata and synonymized a large number of species and subspecies described by Cossmann & Peyrot (1924) with O. excoelata (e.g., O. gibbovaricosa , O. avitensis , O. excoelata var. merignacensis , O. excoelata var. cestasensis ). Ocinebrina excoelata as described by Cossmann & Peyrot (1924) is reminiscent of the Paratethyan species due to its globose last whorl and it bears lirae inside the outer lip, but differs in its much longer, constricted siphonal canal (see Cossmann & Peyrot 1924; pl. 14, figs 35, 38–39, pl. 16, figs 5–6).

Ocinebrinopsis credneri (Hoernes & Auinger, 1885) differs from Ocinebrinopsis gregaria in its much larger size and the coarser spiral sculpture. Ocinebrinopsis dominicii sp. nov. is slenderer and has a higher spire.

Paleoenvironment. Coastal marine, inner neritic environments. The common occurrence at Gainfarn might suggest a preference for seagrass environments (Zuschin et al. 2009).

Distribution in Central Paratethys. Baden (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1995); Vienna Basin: Enzesfeld, Gainfarn, Steinebrunn ( Austria). Pannonian Basin: Herend-Márkó ( Hungary) ( Kókay 1966); Cserhát Mountains ( Hungary) Csepreghy-Meznerics 1954).