Flexopteron goniostoma (Hörnes, 1853)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 123-126

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FF9C-8027-FF50-AE11F90FBC41
treatment provided by	Plazi
scientific name	Flexopteron goniostoma (Hörnes, 1853)
status

Treatment

Flexopteron goniostoma (Hörnes, 1853) View in CoL

Figs 52 View FIGURE 52 , 61A–D View FIGURE 61

[ Murex View in CoL ] goniostomus [sic] Partsch— Hörnes 1848: 18 [nomen nudum].

* Murex View in CoL goniostomus [sic] Partsch—Hörnes 1853: 227, pl. 23, fig. 11.

Murex (Trophon) goniostomus [sic] Partsch—Hoernes & Auinger 1885: 215.

Trophon goniostomus [sic] Ptsch.— Boettger 1902: 32.

Trophon goniostomus [sic] Ptsch. var. citima Bell. — Boettger 1902: 32 [non Flexopteron citima ( Bellardi, 1873) ].

Murex View in CoL (Trophon) goniostomus [sic] Partsch— Schaffer 1908: 105, pl. 11, fig. 5.

Trophon citimus Bellardi — Csepreghy-Meznerics 1956: 400, pl. 5, figs 15–16 [non Flexopteron citima ( Bellardi, 1873) ].

T [rophonopsis]. (P [agodula].) goniostoma (Partsch) View in CoL — Sieber 1958: 145.

Murex View in CoL (Poirieria) goniostoma Partsch View in CoL in Hoernes 1856—Kojumdgieva in Kojumdgieva & Strachimirov 1960: 146, pl. 39, fig. 8, pl. 40, fig. 2.

Trophon citimus Bellardi, 1872 — Strausz 1966: 268, pl. 32, figs 9–10,? pl. 33, fig. 1 [non Flexopteron citima ( Bellardi, 1873) ].

Trophonopsis (Pagodula) goniostoma Partsch — Stancu et al. 1971: 125, pl. 8, fig. 7.

[ Paziella View in CoL ] goniostomus [sic] Partsch in Hörnes, 1853— Vokes 1971: 54.

Trophon citimus Bellardi — Csepreghy-Meznerics 1972: 27, pl. 9, fig. 14 [non Flexopteron citima ( Bellardi, 1873) ].

Trophon citimus (Bellardi, 1872) — Stojaspal 1978: 339 [non Flexopteron citima ( Bellardi, 1873) ].

Trophon goniostomus [sic] (Partsch, 1842)— Stojaspal 1978: 339.

Trophonopsis (Pagodula) goniostoma (Partsch) — Schultz 1998: 65, pl. 26, fig. 1.

Paziella (Flexopteron) goniostoma (Partsch in Hörnes, 1853)— Merle et al. 2011: 170, 522, pl. 139, figs 1–2.

Paziella (Flexopteron) goniostoma (Partsch in Hörnes, 1853)— Kovács et al. 2018: 116, figs 3I–J.

Paziella (Flexopteron) goniostoma (Hörnes, 1853) — Kovács 2019: 118, figs 19–20.

P [aziella]. (F [lexopteron].) citima (Bellardi) — Kovács 2019: 118 [non Flexopteron citima ( Bellardi, 1873) ].

Flexopteron goniostoma (Hörnes, 1853) View in CoL — Kovács & Vicián 2024: 5, figs 2–3.

non Flexopteron cf. goniostoma View in CoL (Partsch in Hoernes, 1853) — Lozouet 2023: 24, pl. 15, figs 1–3).

Type material. Lectotype (designated herein): NHMW 1851 View Materials /0006/0008, SL: 29.1 mm, MD: 17.0 mm, Bad Vöslau ( Austria), illustrated in Hörnes (1853: pl. 23, fig. 11), Figs 61C View FIGURE 61 1 –C View FIGURE 1 3 View FIGURE 3 . Paralectotypes: 12 spec., NHMW 1859 View Materials /0027/0086, Bad Vöslau ( Austria) ;

Illustrated material. NHMW 1866/0001/0644, SL: 28.8 mm, MD: 16.8 mm, Baden-Sooss ( Austria), Figs 61A View FIGURE 61 1 –A View FIGURE 1 3 View FIGURE 3 . NHMW 1866/0001/0644, SL: 25.8 mm, MD: 16.9 mm, Lăpugiu de Sus ( Romania), Figs 61B View FIGURE 61 1 –B View FIGURE 1 3 View FIGURE 3 . NHMW 1875/0009/0023a, SL: 27.1 mm, MD: 15.6 mm, Lăpugiu de Sus ( Romania), Figs 61D View FIGURE 61 1 –D View FIGURE 1 3 View FIGURE 3 .

Additional material. 1 spec., NHMW 1937 View Materials /0002/0288, Bad Vöslau ( Austria) , illustrated in Schaffer (1908: pl. 11, fig. 5); 14 spec., NHMW 1846 View Materials /0037/0212, Baden and Möllersdorf ( Austria) ; 24 spec., NHMW 1863 View Materials /0015/1164, Baden-Sooss ( Austria) ; 7 spec., NHMW 1873 View Materials /0026/0023, Lăpugiu de Sus ( Romania) ; 4 spec., NHMW 1861 View Materials /0040/0072, Hrušovany nad Jevišovkou ( Czech Republic) ; 11 spec., NHMW 1858 View Materials /0047/0015, Porzteich at Břeclav ( Czech Republic) .

Revised description. Small to medium-sized, broad biconic shell with gradate spire, apical angle ~58°. Protoconch pointed, of 1.5 smooth, convex whorls with elevated nucleus. Teleoconch of up to six whorls. Suture moderately incised. Early teleoconch whorls with shallow subsutural ramp, rounded shoulder, subcylindrical below. Axial sculpture of about nine widely spaced, narrow, lamellar varices, forming very short spines along shoulder. Spiral sculpture: first whorl, indistinct cords; second whorl, P1 cord poorly distinct, P1 spine distinct; third whorl, appearance of poorly distinct P2 cord; fourth and fifth poorly distinct P3 above the adapical suture. Last whorl attaining ~70–72% of total height, with broad, shallow, flattened subsutural ramp, angled shoulder, subcylindrical to weakly convex below, strongly constricted at base; up to seven lamellar varices; terminal varix slightly flaring, P1 forming short, adapically curved spines along shoulder. P1, P3, P4, P5 slightly more prominent, P2 atrophied; secondary cords subobsolete, except for weak s2; fasciole weakly swollen, delimiting moderately wide pseudoumbilicus. Aperture moderately wide, ovate-subtrigonal; outer lip thickened, flared in basal area, strongly constricted at siphonal canal, with everted edge; P1 developing long, trigonal spine at shoulder, with groove extending from inner edge of outer lip to P1; prominent D1–D6 at inner edge of lip, D1 stronger. Siphonal canal moderately long, moderately wide, open, straight or slightly deflected to the left, weakly dorsally recurved. Columella smooth, moderately excavated in adapical part, twisted at transition to siphonal canal. Columellar callus forming broad, adherent rim, parietal denticle broad, low or absent, no anal denticle.

Discussion. Bellardi (1873) separated his Murex citimus , from the Burdigalian of Baldissero ( Italy), from Flexopteron goniostoma (Hörnes, 1853) based mainly on its broader shell, lower spire and on the shorter spine on P1. These features, however, are variable in Flexopteron goniostoma and do not allow a reliable distinction. The syntype of Flexopteron citima illustrated in Ferrero Mortara et al. (1981: 37, pl. 2, figs 6a–b) is almost indistinguishable from the Paratethyan specimens except for the more prominent primary cords and subequal secondary cords on the terminal varix. We are not aware of any other specimens of F. citima and its variability is unclear. Provisionally, we keep both species as valid but note that F. goniostoma would have priority over F. citima . The specimen from the Chattian of Saint-Etienn-d’Orthe ( France), illustrated by Lozouet (2023, pl. 15, figs 1–3) and attributed by that author to Flexopteron cf. goniostoma , differs from Flexopteron goniostoma in its multispiral protoconch and represents a yet undescribed species.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Carpathian Foreland Basin: Hrušovany nad Jevišovkou ( Czech Republic) (hoc opus); Vienna Basin: Bad Vöslau, Baden, Gainfarn, Möllersdorf, Niederleis ( Austria) Porzteich at Břeclav ( Czech Republic) (Hoernes &Auinger 1885); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Pannonian Basin: Letkés, Szob, Tekeres ( Hungary) ( Strausz 1966; Kovács et al. 2018, Kovács & Vicián 2024); Southern Carpathians: between Dubova and Pojejena ( Romania) ( Stancu et al. 1971); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1885; Kovács 2019); Dacian Basin: Bozuritsa, Opanec, Staropatica, Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Incertae sedis

Genus Pterynotus Swainson, 1833

Type species. Murex pinnatus Swainson, 1822 View in CoL [= Pterynotus alatus ( Röding, 1798) View in CoL ], by monotypy. Present-day, Indo-West Pacific.

NHMW	Naturhistorisches Museum, Wien

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Muricidae
Genus	Flexopteron