Pterynotus granuliferus ( Grateloup, 1833 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 126-128

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FF9B-8025-FF50-AA1CF88ABE49
treatment provided by	Plazi
scientific name	Pterynotus granuliferus ( Grateloup, 1833 )
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Pterynotus granuliferus ( Grateloup, 1833)

Figs 52 View FIGURE 52 , 62A–B View FIGURE 62

* Murex View in CoL granuliferus Nob. — Grateloup 1833: 96.

Murex View in CoL granuliferus Grat. — Grateloup 1845: pl. 30, fig. 17.

Murex granuliferus Grat. —Hörnes 1853: 254, pl. 25, figs 19a–b.

[ Murex View in CoL granuliferus Grat. ] varieta A.— Bellardi 1873: 79, pl. 6, fig. 4.

Murex View in CoL (Chicoreus) granuliferus Grat. —Hoernes & Auinger 1885: 208.

Murex View in CoL (Chicoreus) granuliferus Grat. — Boettger 1906: 44.

Murex View in CoL ( Chicoreus View in CoL ?) granuliferus var. exgranulosa Sacc. — Sacco 1904: 21.

Murex View in CoL ( Inermicosta View in CoL ) granulifer Grateloup— Cossmann & Peyrot 1924: 111, no. 737, pl. 18, fig. 10 [unjustified emendation]. Murex View in CoL granuliferus Grateloup — Strausz 1943: pl. 3, figs 9, 15.

Ch [icoreus]. (Ch [icoreus].) granuliferus (Grat.) — Sieber 1958: 144.

Murex View in CoL (Chicoreus) granuliferus Grateloup 1840 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 145, pl. 39, fig. 7. [ Pterynotus View in CoL ] granuliferus Grateloup, 1833 — Vokes 1971: 55.

Pterynotus granuliferus ( Grateloup, 1833) — Stojaspal 1978: 334.

Purpura (Tritonalia) granulifera (Grateloup, 1840) — Bałuk 1995: 227, pl. 24, figs 1–2.

Pterynotus View in CoL (s.s.) granuliferus ( Grateloup, 1833) — Merle et al. 2011: 121, 416, pl. 86, figs 1–3.

Pterynotus granuliferus ( Grateloup, 1833) — Landau et al. 2013: 148, pl. 21, figs 12–13.

Pterynotus granuliferus (Grateloup) — Vicián et al. 2017: 268, pl. 2, figs 1–2.

Pterynotus granuliferus ( Grateloup, 1833) — Kovács et al. 2018: 114, figs 3E–F.

Pterynotus granuliferus ( Grateloup, 1833) — Kovács & Vicián 2024: 6, figs 4–5.

Type material. Syntypes from the Early Miocene of Dax ( France) and the Middle Miocene of Saubrigues ( France), one syntype illustrated in Grateloup (1845: pl. 30, fig. 17); The Grateloup’s collection is stored in the University of Bordeaux-Talence ( France), but the syntypes were not located ( Merle et al. 2011: 416).

Illustrated material. NHMW 1853/0003/0115, SL: 44.5 mm, MD: 22.8 mm, Gainfarn ( Austria), illustrated in (Hörnes 1853: pl. 25, figs 19a–b), Figs 62A View FIGURE 62 1 –A View FIGURE 1 3 View FIGURE 3 . SL: 58 mm, MD: 32 mm, Letkés ( Hungary), collection Anton Breitenberger (Bad Vöslau, Austria), Figs 62B View FIGURE 62 1 –B View FIGURE 1 3 View FIGURE 3 .

Revised description. Medium-sized, solid, moderately slender fusiform, trivaricate shell with gradate spire; apical angle initially ~75°, later decreasing to ~40°. Protoconch unknown. Teleoconch of up to six whorls. Early teleoconch whorls convex with deeply incised, shallowly undulating suture. Axial sculpture of three trigonal varices and three slightly weaker intervarical ribs. Spiral sculpture of weak IP and prominent P1–P3. Later whorls with weakly concave subsutural ramp and marked shoulder, convex below; P4 appears above suture on fifth whorl. Last whorl attaining ~70% of total height, broad weakly convex subsutural ramp delimited by angular shoulder, convex below, strongly constricted at base. Subsutural ramp with narrow adis, IP and abis and slightly weaker tertiary cord in interspaces. P1 forming short spines on shoulder varices; P2–P5 slightly stronger than P1, P6 on convex part of whorl, weak, equal in strength to P1, ADP and MP moderately prominent, ABP weak; s1–s6, ads, ms adapically and abapically adjoined by tertiary thread. All cords finely granulose. Varices weakly foliaceous; offset by ~23° between whorls; one weaker intervarical rib between varices, almost as strongly developed as varices. Broad fasciole delimiting wide, shallow pseudoumbilicus. Aperture narrowly ovate. Outer lip with prominent ID, weaker D1–D6, strengthening abapically, D3 occasionally bifid. Anal canal broad, moderately well developed, accentuated by moderately prominent parietal denticle and ID. Siphonal canal of medium length, open, straight, slightly deflected to the left. Columella broadly excavated, smooth. Columellar callus forming broad rim; erect over base, adherent in parietal area.

Discussion. Cossmann & Peyrot (1924: 112) separated specimens from the Tortonian of the Po-Basin ( Italy) and those from the Badenian (Langhian) of the Vienna Basin ( Austria) as distinct species. Sacco (1904: 21) proposed Murex granuliferus exgranulosa as name for the Italian specimens. The Paratethyan specimens examined herein and by Kovács et al. (2018) as well as the specimens from the Serravallian of Turkey, described by Landau et al. (2013) display some variability in spire height, distinctness of the shoulder angulation and strength of spiral sculpture and varices. Therefore, we follow Landau et al. (2013) and consider these occurrences to represent a single widely distributed species.

Paleoenvironment. Inner to outer neritic, marine environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Bad Vöslau, Baden, Gainfarn, Steinebrunn ( Austria) (Hoernes & Auinger 1885); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1885); Pannonian Basin: Letkés ( Hungary) ( Kovács et al. 2018); Korytnica Basin: Korytnica ( Bałuk 1995); Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus ( Romania) ((Hoernes & Auinger 1885; Boettger 1906); Dacian Basin: Opanec, Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akpınar, Lale, Seyithasan ( Turkey) ( Landau et al. 2013); Tortonian (Late Miocene): Po Basin: Stazzano, S. Agata ( Italy) ( Bellardi 1873).

Northeastern Atlantic. Langhian (Middle Miocene): Aquitaine Basin: Saubrigues ( France) ( Cossmann & Peyrot 1924).

Subgenus Pterymarchia Houart, 1995 b

Type species. Murex tripterus Born, 1778 View in CoL ; original designation by Houart (1995 b: 127). Present-day, Indo-West Pacific.

NHMW	Naturhistorisches Museum, Wien

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Muricidae
Genus	Pterynotus