Pterynopsis guzhovi, Harzhauser & Landau & Merle, 2025

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 106

Pterynopsis guzhovi nov. nom.

Figs 51D–E

M [urex]. affinis m.— Eichwald 1830: 224 [non Murex affinis Gmelin, 1791 ].

Murex affinis m. — Eichwald 1851: 100, pl. 8, fig. 10 [non Murex affinis Gmelin, 1791 ].

Murex affinis — Eichwald 1852: plate captions, pl. 8, fig. 10 [non Murex affinis Gmelin, 1791 ].

Mur [ex]. affinis m.— Eichwald 1853: 189 [non Murex affinis Gmelin, 1791 ].

Type material. Lectotype (designated herein): Inv. Nr. 3-372-1, SL: 21.0 mm, MD: 10.8 mm, Żukowce (Zhukivtsi) (Ukarine), illustrated in Eichwald (1851: pl. 8, fig. 10), St. Petersburg State University, Paleontological Museum ( Russia), Figs 51D 1 –D 3 . Paralectotype: Inv. Nr. 3-546-1, SL: 18.3 mm, MD: 8.4 mm, Żukowce (Zhukivtsi) (Ukarine), St. Petersburg State University, Paleontological Museum ( Russia), Figs 51E 1 –E 3 .

Type locality. Żukowce (Zhukivtsi) ( Ukraine) .

Age. Middle Miocene, late Badenian (Serravallian).

Etymology. In honor of Aleksandr Guzhov of the Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, who greatly helped with the identification of Eichwald’s material.

Diagnosis. Slender to moderately slender fusiform, trivaricate shell with up to three intervarical ribs, distinct shoulder and prominent P1–P3 on antepenultimate and penultimate whorls and P1–P4 on last whorl; aperture with thick, flared outer lip and three weak denticles. Varices prominent, not flaring.

Description. Small, slender to moderately slender fusiform, trivaricate shell; apical angle ~52–59°. Protoconch unknown. Teleoconch of up to seven teleoconch whorls. Suture moderately incised, shallowly undulating. Early teleoconch whorls convex with moderately broad subsutural ramp and distinct shoulder. Axial sculpture of three varices and up to three broad intervarical ribs. Spiral sculpture on spire whorls of prominent P1 along shoulder varix and prominent P2 and P3. Last whorl attaining ~67–72% of total height, moderately convex, with rounded shoulder, distinctly constricted below. Three prominent, rounded varices. Prominent P1, P2, P3 and P4. P5, P6 missing; no secondary cords. Aperture subtrigonal. Outer lip edge flared and everted. Denticles reduced (D1–D3) to indistinct swellings placed at inner edge of everted portion of outer lip. Anal canal indistinctly incised; siphonal canal very long, narrow, open, straight, weakly dorsally recurved. Columella weakly excavated in adapical part, smooth, twisted at siphonal canal. Columellar callus forming very broad, adherent rim.

Discussion. Murex affinis Eichwald, 1830 is considered as a junior synonym of Pterynopsis badensis ( Nyst, 1881) in MolluscaBase eds. (2024b). However, the type material of Pterynopsis guzhovi sp. nov. differs from that species in its coarser primary cords (P1 to P3) on the spire whorls, the prominent P2, which is weak in P. badensis , and the prominent intervarical ribs, which are obsolete in P. badensis . Its sculpture on the penultimate whorl resembles that of Pterynopsis meridionalis Lozouet, 1999 (see Lozouet 2023: pl. 32, figs 19–21) from the Chattian of the Aquitaine Basin, but P. meridionalis differs in prominent P5, P6, ADP and MP. Pterynopsis subcontabulata (Millet, 1854) from the Tortonian to the Pleistocene of France and England displays weaker intervarical ribs, reduced P2–P3 and more prominent internal denticles D1–D4 (see Landau et al. 2019, pl. 29). The binome Murex affinis is preoccupied by Murex affinis Gmelin, 1791 and Murex affinis Risso, 1826 . Therefore, we propose Pterynopsis guzhovi as replacement name.

Paleoenvironment. Shallow marine, inner neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Ukrainian Fore-Carpathian Basin: Bilka, Żukowce (Zhukivtsi), Stary Poczajów, Zaleśce (Salisze), Sawadynze ( Ukraine) ( Eichwald 1830, 1853).

Figures

FIGURE 1. Cord diagrams and apertural features for Haustrinae [Haustrum? pyrulatum (Hoernes & Auinger, 1882)] and Muricinae [Bolinus gapus (De Gregorio, 1885a), Bolinus partschi (Hörnes, 1853), Chicoreus (Triplex) aquitanicus (Grateloup, 1833), Chicoreus (Triplex) hirmetzli (Kovács, 2018), Chicoreus (Triplex) notatus (Eichwald, 1830). Hexaplex (Trunculariopsis) trunculoides (Pusch, 1837), Hexaplex (Trunculariopsis) esplus (De Gregorio, 1885a), Hexaplex(Trunculariopsis) subasperrimus (d’Orbigny, 1852), Promurex spinicosta (Bronn, 1831), Exquisitor graniferus (Michelotti, 1841)].

FIGURE 3. Bolinus gapus (De Gregorio, 1885a). A1–A3. NHMW 1847/0046/0030, Szob (Hungary). B1–B3. paralectotype, NHMW 1860/0015/1270, Grund (Austria). C1–C3. NHMW1869/0001/0280, Grund (Austria). D1–D3. lectotype, NHMW 1856/0050/0122, Gainfarn (Austria).

FIGURE 51. A–C. Pterynopsis badensis (Nyst, 1881). A1–A3. holotype, NHMW 1846/0037/0208, Steinebrunn (Austria). B1–B3. NHMW 1860/0040/0143a, Lăpugiu de Sus (Romania). C. Gainfarn (Austria), collection Anton Breitenberger. D–E. Pterynopsis guzhovi nov. nom. D1–D3. lectotype, Inv. Nr. 3-372-1, Żukowce (Zhukivtsi) (Ukraine). E1–E3. paralectotype, Inv. Nr. 3-546-1, Żukowce (Zhukivtsi) (Ukaaine).