Pagodula perditus ( De Gregorio 1885b )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 110

publication ID	https://doi.org/10.11646/zootaxa.5572.1.1
publication LSID	lsid:zoobank.org:pub:7AF6A560-3FAC-4490-B167-327A7912F242
persistent identifier	https://treatment.plazi.org/id/F73C87F9-FF8B-8037-FF50-A9ECFB5DB8B4
treatment provided by	Plazi
scientific name	Pagodula perditus ( De Gregorio 1885b )
status

Treatment

Pagodula perditus ( De Gregorio 1885b)

Figs 52 View FIGURE 52 , 54A–C View FIGURE 54

[ Murex View in CoL ] Klipsteinii Hörnes— Hörnes 1848: 18 [nomen nudum].

Murex varicosissimus Bon. View in CoL —Hörnes 1853: 225, pl. 23, fig. 9 [non Pagodula varicosissima ( Michelotti, 1841) View in CoL ].

Murex (Trophon) varicosissimus Bonelli View in CoL —Hoernes & Auinger 1885: 215 [non Pagodula varicosissima ( Michelotti, 1841) View in CoL ].

* [ Trophon (Trophon) varicosissimus (Bon.) Mich. View in CoL ] F. a perditus — De Gregorio 1885b: 31 [pro Murex varicosissimus sensu Hörnes 1853: 225 View in CoL , pl. 23, fig. 9].

Trophon varicosissimus (Bon.) View in CoL — Boettger 1902: 32 [non Pagodula varicosissima ( Michelotti, 1841) View in CoL ].

T [rophonopsis]. (P [agodula].) varicosissima (Bon.) View in CoL — Sieber 1958: 145 [non Pagodula varicosissima ( Michelotti, 1841) View in CoL ].

Trophon varicosissimus Bonelli, 1826 View in CoL — Strausz 1966: 269, pl. 32, figs 5–6 [non Pagodula varicosissima ( Michelotti, 1841) View in CoL ].

Trophon varicosissimus (Bonelli, 1826) View in CoL — Stojaspal 1978: 339 [non Pagodula varicosissima ( Michelotti, 1841) View in CoL ].

non Murex varicosissimus Bon. View in CoL — Simionescu & Barbu 1940: 125, pl. 2, fig. 25 [= Ocinebrina striata ( Eichwald, 1830) View in CoL ].

non Poirieria (Pagodula) varicosissima ( Michelotti, 1841) — Kovács 2019: 117, figs 17–18 [= Pagodula spinifera ( Csepreghy-Meznerics, 1969) ].

Type material. Holotype, NHMW 1869 View Materials /0001/0172, SL: 25.4 mm, MD: 14.3 mm, Baden ( Austria), illustrated in Hörnes (1853: pl. 23, fig. 9), Figs 54C View FIGURE 54 1 –C View FIGURE 1 3 View FIGURE 3 . Trophon (Trophon) varicosissimus perditus De Gregorio, 1885b was introduced for the specimen illustrated in Hörnes (1853: pl. 23, fig. 9).

Illustrated material. NHMW X1866 /0001/0643, SL: 25.4 mm, MD: 12.9 mm, Baden-Sooss ( Austria), Figs 54A View FIGURE 54 1 –A View FIGURE 1 3 View FIGURE 3 . NHMW 2024 View Materials /0047/0003, SL: 27.1 mm, MD: 13.5 mm, Bad Vöslau ( Austria), Figs 54B View FIGURE 54 1 –B View FIGURE 1 3 View FIGURE 3 .

Additional material. 1 spec., NHMW 1869 View Materials /0001/0034, Bad Vöslau ( Austria) ; 3 spec., NHMW 1869 View Materials /0001/0570, Steinebrunn ( Austria) ; 7 spec., NHMW 1869 View Materials /0001/0082, Baden-Sooss ( Austria) ; 3 spec., NHMW 1867 View Materials /0019/0088, Coşteiu de Sus , ( Romania) .

Description. Medium sized, fusiform shell with conical spire; apical angle ~52°. Protoconch unknown. Teleoconch of up to six whorls. Early teleoconch whorls with weakly convex subsutural ramp, weak mid-whorl angulation, slightly constricted below. Axial sculpture of about ten, trigonal ribs. No spiral sculpture. Short P1 spines present on the early whorls. Later teleoconch whorls with broad, weakly concave subsutural ramp; weakly laminar varices forming indistinct, open spines along shoulder. Last whorl attaining ~70% of total height, with broad subsutural ramp, angled to slightly rounded shoulder, weakly convex below, abruptly constricted at base. About ten laminar varices, P1 forming short triangular spines along shoulder; fasciole weak, separated by narrow chink from siphonal canal. P2, P3 and P4 obsolete. Aperture pyriform; outer lip smooth or with five weak denticles placed some distance behind peristome and corresponding to D1 to D4 (D2 bifid); anal canal subobsolete; siphonal canal long, moderately narrow, slightly deflected to the left, shallowly notched. Columella moderately to weakly excavated, twisted at siphonal fasciole. Columellar callus forming narrow adherent rim, poorly delimited from base.

Discussion. The Paratethyan Pagodula perditus ( De Gregorio 1885b) was identified by most authors as Pagodula varicosissima ( Michelotti, 1841) from the Tortonian of Tortona ( Italy). Michelotti (1841) depicted a small biconic shell with an apical angle of about 55°, a steep, broad subsutural ramp and the shoulder placed close above the abapical suture. The type specimen of Pagodula varicosissima seems to have been already lost in the 19 th century because Bellardi (1873) and Sacco (1904) did not re-illustrate the specimen. Pagodula varicosissima from the Tortonian of S. Agata and Montegibbio ( Italy), illustrated by Sacco (1904: pl. 7, figs 16–17) and Montanaro (1935: pl. 4, figs 8–9), differs distinctly in its slenderer shape and much higher spire. Thus, the status of Pagodula varicosissima is unclear. Michelotti’s Pagodula varicosissima differs from Pagodula perditus in its biconic outline and lacks the subcylindrical part of the last whorl of P. perditus . Pagodula varicosissima sensu Sacco (1904) and Montanaro (1935) is clearly much slenderer. Note that the specimen from the Tortonian of Stazzano ( Italy) illustrated by Caprotti (2011: figs 6Q–R) is a turrid ( Brunetti et al. 2012).

Pagodula perditus differs from Pagodula spinifera ( Csepreghy-Meznerics, 1969) in its stockier shape, less gradate spire, more convex last whorl, much weaker spines and broader siphonal canal.

Paleoenvironment. Middle to outer neritic, marine environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Voronyaky Hills: Hołubica (Holubytsia) ( Ukraine) (Hoernes & Auinger 1885); Carpathian Foreland Basin: Rudice ( Czech Republic) (Hoernes & Auinger 1885); Vienna Basin: Baden, Baden Sooss, Bad Vöslau, Enzesfeld, Steinebrunn ( Austria) (Hoernes & Auinger 1885); Eisenstadt-Sopron Basin: Sopron ( Hungary) ( Strausz 1966).