Pterynopsis badensis ( Nyst, 1881 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 104-105

Pterynopsis badensis ( Nyst, 1881)

Figs 47 , 51A–C

M [urex]. tortuosus Sow. —Pusch 1836: 519 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

M [urex]. tripteris Lam.—Pusch 1836: 519 [non Murex tripterus Born, 1778 = Pterynotus tripterus ( Born, 1778) ].

Murex tripteris Lam.—Pusch 1837: 137 [non Murex tripterus Born, 1778 = Pterynotus tripterus ( Born, 1778) ].

Murex tortuosus Sow. —Pusch 1837: 137 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

[ Murex ] affinis Eichw. — Hauer 1837: 418 [non Murex affinis Gmelin, 1791 ].

Murex tortuosus Sow. —Hörnes 1853: 249, pl. 25, figs 12a–c [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

* M [urex]. badensis — Nyst 1881: 4 [nov nom. pro Murex tortuosus sensu Hörnes, 1856 , pl. 25, fig. 12 non J. Sowerby, 1823].

Murex affinis —Quenstedt 1884: 648, pl. 211, fig. 15 [non Murex affinis Gmelin, 1791 ].

Murex (Pteronotus) tortuosus Sow. —Hoernes & Auinger 1885: 203 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Murex tortuosus Sow. — Friedberg 1912: 164, pl. 10, figs 1–2 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Murex tortuosus Sow. — Friedberg 1938: 115 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Murex ( Pterynotus , sect. Pterynotus ) tortuosus Sow. — Korobkov 1955: plate captions, pl. 70, figs 7a–b, pl. 73, figs 8a–b [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Murex tortuosus Sowerby, 1823 — Moisescu 1955b: 141: pl. 12, figs 9–10 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

P [terynotus]. (P [terochelus].) tortuosus (Sow.) — Sieber 1958: 145 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Murex tortuosus Sowerby, 1823 — Zelinskaya et al. 1968: 186, pl. 44, figs 11–12 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Murex badensis Nyst, P.H., 1881 — Vokes 1972: 5.

Pterynotus binominatus (Staat in Cossmann, 1909) — Stojaspal 1978: 334 [non Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Pterynotus (Pterynotus) tortuosus (J. Sowerby, 1823) — Bałuk 2006: 207, pl. 11, fig. 6 [non Murex tortuosus J. Sowerby, 1823 = Pterynopsis binominatus (Staadt in Cossmann, 1909)].

Pterynopsis badensis ( Nyst, 1881) — Biskupič 2016: 39, fig. 1.

Type material. Holotype: NHMW 1846 /0037/0208, SL: 58.6 mm, MD: 34.0 mm, Steinebrunn ( Austria), illustrated in Hörnes (1853, pl. 25, figs 12), Figs 51A 1 –A 3 .

Illustrated material. NHMW 1860/0040/0143a, SL: 61.0 mm, MD: 31.1 mm, Lăpugiu de Sus ( Romania), Figs 51B 1 –B 3 . SL: 53 mm, MD: 28 mm, Gainfarn ( Austria), collection Anton Breitenberger (Bad Vöslau, Austria), Figs 51C .

Additional material. 8spec., NHMW1871 /0010/0347, Steinebrunn ( Austria) ; 2spec., NHMW1846 /0037/0208a, Gainfarn ( Austria) ; 1 spec. NHMW 1866 /0040/0194, Baden-Sooss ( Austria) ; 5 spec., NHMW 1861 /0001/0436, Vienna / Grinzing ( Austria) ; 4 spec., NHMW 1866 /0040/0119, Marz ( Austria) ; 2 spec., NHMW 1863 /0015/0671, Niederleis ( Austria) .

Revised description. Very large, slender fusiform, trivaricate shell; apical angle ~40–50°. Protoconch unknown. Teleoconch of up to seven teleoconch whorls. Suture moderately incised, shallowly undulating. Early teleoconch whorls convex with steep, convex subsutural ramp and weak shoulder. Axial sculpture of three sharp varices and up to two broad, weak intervarical ribs. Spiral sculpture subobsolete. P1 forming short, adapically curved spines along shoulder varix. IP and P2 only faintly developed on varices. Last whorl attaining ~70% of total height, moderately convex, with rounded shoulder, distinctly constricted below. Three flaring varices aligned axially along shell, slightly offset by ~23° between whorls. P1 forming short, broad spine; P2 reduced, P3 and P4 with shorter, less distinct spines. P5, P6, ADP and MP forming low, broad cords; s2–s4, s6 weak to subosolete. Aperture subtrigonal. Outer lip edge flared and everted with weak incision at shoulder-spine. Denticles reduced (D1–D3) to indistinct (D4–D6) swellings placed at inner edge of everted portion of outer lip. Anal canal indistinctly incised; siphonal canal very long, narrow, open, adapical portion slightly bent to the left, abapical portion curved to the right, weakly dorsally recurved. Columella weakly excavated in adapical part, smooth, twisted at siphonal canal. Columellar callus forming very broad, adherent rim.

Discussion. Most Paratethyan authors have identified this species as Murex tortuosus J. Sowerby, 1823 [= Pterynopsis binominatus (Staadt in Cossmann, 1909)] from the Pliocene North Sea Basin. However, the Paratethyan specimens differ in being somewhat smaller (maximum height 44 mm; fide Marquet, 1997, p. 88), higher spired and having spines developed at P1 and P3 on the terminal varix. Moreover, the siphonal canal is shorter and although also bent to the left, lacks the distal bend to the right seen in Pterynopsis badensis ( Nyst, 1881) . Pterynopsis subcontabulata (Millet, 1854) , from the Tortonian to Pleistocene of the northeastern Atlantic, is highly reminiscent of Pterynopsis badensis and differs mainly in its much smaller size (maximum height 27.2 mm; fide Ceulemans et al., 2016, p. 43), prominent denticles in the outer lip and the comparatively straighter siphonal canal (see Ceulemans et al. 2016: pl. 2, figs 2–4). For further discussion on European Pliocene congeners, see Ceulemans et al., 2016, p. 43).

Paleoenvironment. Known from inner neritic to outer neritic, marine environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Faszczówka, Hołdy, Poczajów Stary, Ternopol (Ternopil), Zborów (Zboriv), Żukowce (Zhukivtsi), Zaleśce (Salisze) ( Friedberg 1912, 1938); Voronyaky Hills: Hołubica (Holubytsia), Podhorce (Pidhirtsi) ( Ukraine) ( Friedberg 1912, 1938); Korytnica Basin: Korytnica ( Bałuk 1995); Vienna Basin: Baden-Sooss, Gainfarn, Niederleis, Steinebrunn, Vienna /Grinzing ( Austria) (Hoernes & Auinger 1885), Devínska Nová Ves, Rohožník-Konopiská, ( Slovakia) ( Schaffer 1898; Biskupič 2016); Eisenstadt-Sopron Basin: Marz ( Austria); Făget Basin: Buitur, Lăpugiu de Sus ( Romania) ( Moisescu 1955b; hoc opus).

Figures

FIGURE 1. Cord diagrams and apertural features for Haustrinae [Haustrum? pyrulatum (Hoernes & Auinger, 1882)] and Muricinae [Bolinus gapus (De Gregorio, 1885a), Bolinus partschi (Hörnes, 1853), Chicoreus (Triplex) aquitanicus (Grateloup, 1833), Chicoreus (Triplex) hirmetzli (Kovács, 2018), Chicoreus (Triplex) notatus (Eichwald, 1830). Hexaplex (Trunculariopsis) trunculoides (Pusch, 1837), Hexaplex (Trunculariopsis) esplus (De Gregorio, 1885a), Hexaplex(Trunculariopsis) subasperrimus (d’Orbigny, 1852), Promurex spinicosta (Bronn, 1831), Exquisitor graniferus (Michelotti, 1841)].

FIGURE 3. Bolinus gapus (De Gregorio, 1885a). A1–A3. NHMW 1847/0046/0030, Szob (Hungary). B1–B3. paralectotype, NHMW 1860/0015/1270, Grund (Austria). C1–C3. NHMW1869/0001/0280, Grund (Austria). D1–D3. lectotype, NHMW 1856/0050/0122, Gainfarn (Austria).

FIGURE 47. Cord diagrams and apertural features of Ocenebrinae [Jaton ampistus (De Gregorio, 1885a), Jaton sowerbyi (Michelotti, 1841), Pteropurpura polonica (Pusch, 1836), Pterynopsis badensis (Nyst, 1881)].

FIGURE 51. A–C. Pterynopsis badensis (Nyst, 1881). A1–A3. holotype, NHMW 1846/0037/0208, Steinebrunn (Austria). B1–B3. NHMW 1860/0040/0143a, Lăpugiu de Sus (Romania). C. Gainfarn (Austria), collection Anton Breitenberger. D–E. Pterynopsis guzhovi nov. nom. D1–D3. lectotype, Inv. Nr. 3-372-1, Żukowce (Zhukivtsi) (Ukraine). E1–E3. paralectotype, Inv. Nr. 3-546-1, Żukowce (Zhukivtsi) (Ukaaine).