Timbellus perlongus ( Bellardi, 1873 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Central Paratethys Sea (Haustrinae, Muricinae, Ocenebrinae, Pagodulinae, Typhinae, Muricidae incertae sedis), Zootaxa 5572 (1), pp. 1-162 : 135

Timbellus perlongus ( Bellardi, 1873)

Figs 64 , 67A–B

* Murex perlongus Bell. — Bellardi 1873: 75, pl. 5, figs 8a–b.

Murex (Chicoreus) perlongus Bell. —Hoernes & Auinger 1885: 207, pl. 24, fig. 13.

Murex ( Pteropurpura ) lamarcki [sic] Grateloup— Cossmann & Peyrot 1924: 103, n°730, pl. 13, figs 18–20, pl. 18, fig. 5 [non Timbellus lamarckii ( Grateloup, 1845) ].

[ Pterynotus ] perlongus Bellardi, 1872 [sic]— Vokes 1971: 81.

Pterynotus perlongus (Bellardi, 1872 [sic]) — Stojaspal 1978: 334.

Pterynotus (Pterynotus) perlongus ( Bellardi, 1873) — Bałuk 1995: 215, pl. 24, fig. 8.

Pterynotus perlongus (Bellardi, 1872 [sic]) — Lozouet et al. 2001: 54, pl. 24, figs 2a–b.

Timbellus perlongus (Bellardi, 1872 [sic])— Merle et al. 2011: 454, pl. 105, figs 1, 4, 6.

Timbellus perlongus ( Bellardi, 1873) — Kovács et al. 2018: 114.

Timbellus perlongus ( Bellardi, 1873) — Kovács 2019: 117, figs 11–12.

Timbellus perlongus ( Bellardi, 1873) — Lozouet 2023: 26, pl. 22, figs 11–12.

Type material. The syntypes from the Burdigalian of Termô-Fôurà ( Italy) were stored in the Rovasenda collection, which was transferred to the Museum of Geology and Paleontology in Torino ( Ferrero Mortara et al. 1981: 17). The specimens are lost ( Merle et al. 2011: 454) because a large part of this collection was destroyed ( Merlino & Campanino (2002: 8).

Illustrated material. NHMW 1846/0037/0209, SL: 73.1 mm, MD: 30.1 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1884: pl. 24, fig. 13), Figs 67B 1 –B 2 . NHMW 1857/0014/0017, SL: 76.5 mm, MD: 31.1 mm, Lăpugiu de Sus ( Romania), Figs 67A 1 –A 2 .

Revised description. Large, solid, slender fusiform, trivaricate shell with high spire; apical angle ~40°. Protoconch not preserved in Paratethyan material; conical of two smooth whorls with faint angulation above abapical suture based on specimens from France ( Lozouet 2023: pl. 22, figs 11–12). Teleoconch of up to seven whorls. Suture strongly undulating, moderately incised. Early teleoconch whorls convex with three narrow, strongly raised varices, starting on second teleoconch whorl, one intervarical knob placed mid-whorl. Spiral sculpture largely reduced except for subosolete P1 and P2 on late spire whorls. Varices axially aligned, offset by about 22° between whorls. Last whorl attaining ~68% of total height, moderately convex with indistinct shoulder, weakly constricted. Three foliaceous varices and one weak intervarical rib forming weak knobs at intersections with P1–P3. Spiral sculpture of broad, low P1–P6, ADP, MP,ABP, developed mainly on varices. No secondary cords. Terminal varix slightly flaring. Fasciole moderately swollen, delimiting moderately broad, shallow pseudoumbilicus. Aperture narrowly ovate; outer lip thickened, with weakly crenulate edge, with prominent D1–D6 placed some distance behind peristome, D4 bifid; interspaces between denticles abapically narrowing. Anal canal broadly U-shaped; siphonal canal long, open but extremely narrow, straight, with very broad terminal varix. Columella straight, smooth. Columellar callus forming broad rim, sharply delimited, erect at siphonal canal, adherent and expanding in parietal area.

Discussion. Lozouet et al. (2001) and Merle et al. (2011) reported the Proto-Mediterranean Timbellus perlongus ( Bellardi, 1873) also from the northeastern Atlantic.The Middle Miocene Paratethyan specimens are stratigraphically slightly younger and have a slightly higher spire but sculpture and apertural features agree fully with the Early Miocene specimens. Timbellus perlongus differs from other Paratethyan congeners in its large-sized, thicker shell with a somewhat distorted outline.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1995); Pannonian Basin: Letkés ( Hungary) ( Kovács et al. 2018); Făget Basin: Lăpugiu de Sus ( Romania) ( Kovács 2019).

Proto-Mediterranean Sea. Burdigalian (Early Miocene): Colli Torinesi: Termô-Fôurà ( Italy) ( Bellardi 1873).

Northeastern Atlantic. Aquitanian/Burdigalian (Early Miocene): Aquitaine Basin: Léognan, Meilhan, Mérignac, Saint-Martin d’Oney, Saint-Paul-lès-Dax, Saucats ( France) ( Lozouet et al. 2001; Lozouet 2023).

Figures

FIGURE 1. Cord diagrams and apertural features for Haustrinae [Haustrum? pyrulatum (Hoernes & Auinger, 1882)] and Muricinae [Bolinus gapus (De Gregorio, 1885a), Bolinus partschi (Hörnes, 1853), Chicoreus (Triplex) aquitanicus (Grateloup, 1833), Chicoreus (Triplex) hirmetzli (Kovács, 2018), Chicoreus (Triplex) notatus (Eichwald, 1830). Hexaplex (Trunculariopsis) trunculoides (Pusch, 1837), Hexaplex (Trunculariopsis) esplus (De Gregorio, 1885a), Hexaplex(Trunculariopsis) subasperrimus (d’Orbigny, 1852), Promurex spinicosta (Bronn, 1831), Exquisitor graniferus (Michelotti, 1841)].

FIGURE 2. A1–A2. Haustrum? pyrulatum (Hoernes & Auinger, 1882), holotype, NHMW 1859/0045/0590, Lăpugiu de Sus (Romania). B1–B2. Haustrum lacunosum (Bruguière, 1789), MNHN-IM-2000-904, lectotype, New Zealand, present-day, photo: Manuel Caballer, 2014 RECOLNAT project ANR-11-INBS-0004.

FIGURE 64. Cord diagrams and apertural features of Muricidae incertae sedis [Timbellus latifolius (Bellardi, 1873), Timbellus kovacsi sp. nov., Timbellus perlongus (Bellardi, 1873), Timbellus trinodosus (Bellardi, 1873), Timbellus weinmannae sp. nov., Purpurellus cyclopterus (Millet, 1865), Phyllocoma sp., Vitularia linguabovis (de Basterot, 1825)].

FIGURE 67. Timbellus perlongus (Bellardi, 1873).A1–A2. NHMW 1857/0014/0017, Lăpugiu de Sus (Romania). B1–B2.NHMW 1846/0037/0209, Lăpugiu de Sus (Romania).