Chrysorthenches, DUGALE, 1996

Sohn, Jae-Cheon, Kobayashi, Shigeki & Yoshiyasu, Yutaka, 2020, Beyond Wallace: a new lineage of Chrysorthenches (Lepidoptera: Yponomeutoidea: Glyphipterigidae) reveals a journey tracking its host-plants, Podocarpus (Pinopsida: Podocarpaceae), Zoological Journal of the Linnean Society 190, pp. 709-736 : 712

GENUS CHRYSORTHENCHES DUGALE, 1996

Chrysorthenches Dugdale, 1996: 34 . Type species: Orthenches porphyritis Meyrick, 1886 , by original designation.

Dugdale (1996) defined Chrysorthenches in relation to Orthenches and suggested the following apomorphies for the former genus: (1) in the male genitalia, the gnathos and socii absent; (2) the anellus as a spinulose sheath and often with strong apical thorns; (3) the vinculum plus saccus T-shaped; (4) the posterior margin of the saccus convex; (5) the male phallus uniformly cylindrical; (6) the male sternum VIII with a V-shaped lobe ( Fig. 2G); (7) the male tergopleural lobe arising obliquely and fused dorsally ( Fig. 2G, arrow); and (8) the larval abdominal segment VII with spiracle and seta SD1 on a common pinaculum or scobinate field.

Two additional characters: the presence of the antennal scape with pecten, but no awning, and the valva of the male genitalia with an outer tuft of specialized scales or setae distally, were suggested as apomorphies for Chrysorthenches by Dugdale (1996). These were excluded from this study as proposed from our cladistics analysis. With our additions, Chrysorthenches comprises 13 species in three species-groups.