Pinachenium longicorne ( Baudi di Selve, 1848 )

Struyve, Tim, 2024, Revision of the Afrotropical Pinophilina (Coleoptera: Staphylinidae: Paederinae) Part 1: genera, Pinachenium, Pinoplanus, Pinobelus and Pinocolis, Belgian Journal of Entomology 152, pp. 1-75 : 11-12

Pinachenium longicorne ( Baudi di Selve, 1848)

Figs 2 C-D, 3 C

Pinophilus longicorne BAUDI DI SELVE, 1848: 139

Pinophilus depressus KRAATZ, 1859: 157 , synonymized by ASSING, 2022: 94

Pinophilus planus KRAATZ, 1859: 157 , synonymized by ASSING, 2022: 94

Pinophilus verticalis FAUVEL, 1895: 219 , synonymized by ASSING, 2022: 94

Pinophilus wittmeri Koch, 1934: 70 , synonymized by ASSING, 2022: 94

TYPE MATERIAL EXAMINED

Egypt: Kirdassah, 24/IX/33, W. Wittmer: cotypus P. wittmeri : 2 ind. ( MNHN) .

NON TYPE MATERIAL EXAMINED. Angola: Rocadas , R. Cunene, 19-22/II/1972: 2 ind. ( NHMUK) ; Botswana: Maun, Island Safari Lodge, 2-15/I/1994, M. Snižek leg.: 1 ind (cJan); Chobe distr. Kasane, 1/I/1994, Becnačik leg.: 1 ind. (cJan); bor. 1/1/1994, Kasane, M. Snižek lgt.: 3 ind. (cJan); bor. Maun, 2-15/I/1994, Island Safari Lodge env. M. Snižek lgt .: 1 ♂ (cJan); bor. Maun , 2-15/I/1994, Island Safari Lodge env. M. Snižek lgt .: 1 ♂ (cJan); Chad: Salamat, Zakouma village, VII/2014 C. Vanderbergh leg.: 1 ind. (cStr); N’Djemena, 29/07/2006, C. Vanderbergh leg.: 1 ind. (cStr); Congo-Kinshasa : Lac Edguard : Vitshumbi, U.V., 31/III/1953, J. Verbeke leg.: 1 ind. ( RBINS) ; Lac Albert, Mwita prcs foret galerie, U.V., 22/XII/1953, J. Verbeke leg.: 1 ind. ( RBINS) ; Ethiopia: Harer, 30.5 km, N.E. of Dire Dawal , 300 m, 0936N, 4153E, Black light, R.O.S. Clarcki: 1 ind. ( NHMUK) ; Ilubabor Gambela, XI/1972, R.O.S. Clarck, Black light: 1 ind. ( NHMUK) ; Shoa Awash Park 08572N 3957E, 1000 B.M. 1973-450 R.O.S. Clark: 1 ind. ( NHMUK) ; Harer prov. Gode-Ogaden , 7/XI/1974, coll. T.J. Crowe: 1 ind. ( RMCA) ; Ivory Coast: Lamto ( Toumodi ), III/1968, U.V., Girard leg.: 1 ind. ( MNHN) ; Lamto ( Toumodi ), 22/V/1962, U.V.: 1 ind. ( MNHN) ; Lamto ( Toumodi ), 10/VII/1962: 2 ind. ( MNHN) ; Lamto ( Toumodi ), 1-15/XII/1966, lumière, Girard: 1 ind. ( MNHN) ; Lamto ( Toumodi ): 2 ind. ( MNHN) ; Kenya: S. Garissa env. 30/XI/1999, M. Snižek lgt.: 2 ind. (cJan); S. Voi , 13-17/ XII/1997, M. Snižek lgt.: 1 ind. (cJan); S.E. 11/IV/2004, Lamu prov. , E of Garsen, M. Snižek lgt.: 1 ind. (cJan); SE, Tsavo n.p., Sagala hills, Voi env. 23/XI-4/XII/2000, Rudolf Kmeco leg.: 1 ind. (cJan); Madagascar: Mahajanga prov., Mahajanga riv., Ampatika env. Ivo Jeniš leg. 17-19/XI/1995: 1 ind. ( MNHN) ; Tanandava , lumière, 1963-1964, G. Schmitz: 3 ind. ( RMCA) ; Mauritania: 31/VII/1964, U.V., S.L. Amiet: 1 ind. ( MNHN) ; 1964, Amiet leg: 2 ind ( MNHN); Namibia: Kavango, Rundu , Okavango nv. 16-19/I/1999, 1050 m, Rudolf Kmeco leg.: 1 ind. (cJan); East Caprivi, Katima Mulilo, Zambezi riv. 19-25/I/ 1999, 950 m, Rudolf Kmeco leg.: 1 ind. (cJan); bor. or. 15-24/I/1995, Katima Mulilo, Caprovi Zipfel, M. Snižek lgt.: 1 ind. (cJan); ZMB, 1992, East Caprivi: Katima Mulilo, lux, 17°29’S, 24°17’E, 3-8/III/1992, leg. M. Uhlig: 1 ind. ( ZMHB) ; ZMB, 1992, E. Caprivi: 30 km SE, Katima Mulilo , 17°31’S 24°25’E, Zambezi- Altwasserarm, lux, 6/III/1992, lg. M. Uhlig: 2 ind. ( ZMHB) ; 1/III/1994, 18°14’S, 21°43’E, Kavango Mahango Game Reserve, Piknik site, lux, leg. M. Uhlig: 2 ind. ( ZMHB) ; Senegal: Richard-Toll, XI/1967, A. Descarpentries, T. Leye et A. Villiers: 21 ind. ( MNHN, cStr); Piège UV, 1/XII/1960, Richard-Toll: 1 ind. ( MNHN) ; Saint-Louis, 16/IX/59, Ifan , 16/IX/1963, T. Lège: 1 ind. ( MNHN) , Richard-Toll , 12/XII/1963, R. Roy, Piège à rayons ultra-violets: 1 ind. ( MNHN) ; Piège UV, 1/XII/1960, Richard-Toll: 1 ind. ( MNHN) ; Piège UV, 2/XII/1960, Richard-Toll: 1 ind. ( MNHN) ; Botou Village , 13°48’28”N, 13°34’43”W, 18-25/VI/2019, MV light trap, Aristophanous, M., Moretto, P., Mulvaney, L. leg.: 1 ind. ( NHMUK) ; Mt. Bourv, St. Orstom , 2/198 I, B. Sigwalt leg.: 1 ind. ( MNHN) ; Somalia: 22-23/VII/2021, NW of Xoorey , ca 840 m, 10°10’59”N, 43°21’49”E, D. Král & D. Sommer lgt.: 1 ind. ( NMPC) ; 17-18/VII/2021, S. of Habaas, ca. 850 m, 10°24’40”N, 42°48’45”E D. Král & D. Sommer lgt.: 2 ind. ( NMPC) ; South Africa: Limpopo , 10 km E of Stockpoort, at Limpopo riv., 16-19/XII/2006, M. Bednařik leg.: 2 ind. (cJan); Transvaal near Thabazimbi , 15-16/I/1999, leg. P. Schüle: 1 ind. ( SMNS) ; 30/I-3/II/1995, KwaZulu-Natal, N’dumu Game Res. 26°55’S, 32°19’E, leg. F. Koch: 1 ind. ( ZMHB) ; South-Sudan: Upper Nile, Malakal , 5-20/I/1963, Linnavuori, ad lucem: 1 ind. ( NHMW) ; Juba , 27/II-2/III/1963: Linnavuori R. leg., 4.817°N, 31.58°E: 1 ind. ( MZH) ; Sudan: Kharthoum 22/XIII/1966, P. Štys leg.: 4 ind. (cJan); Kharthoum 28/XIII/1966, P. Štys leg.: 1 ind. (cJan); Kharthoum prov. , Shambat , 15/X/1966, P. Štys leg.: 3 ind. (cJan); Shambat , 22/XI/1966, P. Štys leg.: 1 ind. (cJan); Prov. Bleu Nile, Wad Shafie , 20 km W von Wad Medani , am licht, 27/IX/1975, Bremer legit: 2 ind. ( RBINS) ; Prov. Bleu Nile, Wad Medani , am licht, 23/IX/1975, Bremer legit: 2 ind. ( RBINS) ; Prov. Bleu Nile, Wad Medani , am licht, 5/III/1974, Bremer legit: 1 ind. ( RBINS) ; Kosti , 22/I/1963, Linnavuori: 2 ind. ( NHMW) ; Tanzania: pr. Mgorogoro , 6° 59.6°N, 37°15.2’E, 450m, 50 km SW of Mgorogoro, leg. F. Kantner, 12/I/2007: 5 ind. ( SMNS) ; o.mer. 8°03.2’S, 38° 52.7°E, 70 m, 15 km N Utete , pr. Pwani, leg. F. Kantner, 16/I/2007: 1 ind. ( SMNS) ; Morogoro Prov. Wami-DumilaVII, Kilosa Distr 29/XI/2008, G. Mpoyola, under rot logs: 1 ind. ( FMNH) ; Iringa prov. Iringa Tanangozi Area , 13/ 16/XI/2008, K. Angelus ex rot logs dry mushrooms: 1 ind. ( FMNH) ; Zimbabwe: Salisbury, Mashomaland : 1 ind. ( NHMUK) .

COMMENTS

Variable species, with tendency for forming local distinctive populations. There are several forms that could be described as separated (sub)species, but always intermediate forms are found. Thus, the decision to keep it as one species. The western individuals ( Ivory Coast, Mauritania, Senegal) have the micropunctation on the head sparser, while most, but not all, individuals from other locations the micropunctation is more dense (e.g., specimens from Uganda and Zambia with also sparser micropunctation). The color seems also regionally variable: western species are brighter, eastern darker. One individual from Madagascar is completely dark, two other from Madagascar completely reddish, with no other differences.

This species seems widely distributed in the entire sub-Sahara Africa, but also Egypt and Cyprus up to India and Sri Lanka ( ASSING, 2022).

Figures

Fig. 2. A-F, Head, dorsal view. A, Pinachenium abessinum (Bernhauer, 1915). B, P. insolitum (Fagel, 1963). C-D, P. longicorne (Baudi di Selve, 1848). E, P. nitidum sp. nov. F, P. pseudabessinum (Gridelli, 1928). Scale bar: 1 mm.