Bolanthus frankenioides (Boiss.) Barkoudah., Wentia

Yildizbaş, Avni & Koç, Murat, 2025, The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye, Phytotaxa 696 (1), pp. 28-54 : 39-41

Bolanthus frankenioides (Boiss.) Barkoudah., Wentia 9: 169 (1962) ≡ Gypsophila frankenioides Boiss., Diagn. Ser.

1(1): 10 (1843)

Lectotype [designated by Barkoudah (1962: 169) as “ holotype ”, here corrected according to the Art. 9.10 of ICN]: — [ Turkey C2 Denizli] in regione alpina Cariae, Cadmus orientalis supra Khonas [Honaz Da.] ad nives, June 1842, Boissier s. n. ( G00544521 ! [photo]; isolectotype: P01903171 !).

Description:— Perennial herbs. Stems prostrate, 5v50, 8-15 cm, base 0.7–1 mm diameter, loose branched, complately glandular hairs, nodes 3–6, internodes 5–15 mm, glandular hairs. Leaves linear-oblanceolate, 3–5 × 0.5–0.7 mm, glandular hairs, sparsely glandular hairs or glabrous prominent 3-nerved, greenish, not adpressed to stem; terminal acute; sheaths membranous, 0.2–0.3 mm, glandular hairs. Inflorescence in apex and sides, glandular hairs; flowers pedicellate, 3–15-flowers. Bracts linear-oblanceolate, 4–5.5 × 0.5–0.7 mm, almost as much as inflorescence; margin membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Pedicels 1–4 mm, glandular hairs. Calyx tubulate, glandular hairs, 3–3.5 × 0.9–1.1 mm; calyx teeth narrowly triangular, 1–1.3 mm tall, margins ciliate, terminal acute. Petals oblanceolate, 4–5 × 0.6–1 mm, up to 1.5 times sepals, lamina pink, petal petal nerves dark pink; terminal cute; base cuneate. Stamens 10, filament 3–4 mm tall. Styles 2, 3-4 mm tall. Capsule oblong, 2.5–3 × 0.9–1.1 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 1–1.1 × 0.8–0.9 mm, brown, side and ridge surface tuberlculate, granular.

Pedicels 2–4 mm, as long as calyx...................................................................................................................................... var. frankenioides Pedicels 1–2 mm, shorter than calyx ..................................................................................................................................... var. fasciculatus

Typification of the name Gypsophila frakenioides :— The protologue of Gypsophila frankenioides ( Boissier 1843: 10) consists of a detailed diagnosis and the provenance (“Hab. in pratis ad nives deliquescentes in parte superiori montis Cadmi supra Colossam ubi legi Junio 1842 ”). Barkoudah (1962: 169) stated that the holotype and isotype are at G and P, respectively. Also Madhani et al. (2024: 1245) indicated a holotype at G (G00544521), as well as a possible isotype (G00544571). However, no holotype was explicitly reported in the protologue by Boissier (1843: 10). By checking the herbarium G, we noted that Barkoudah annotated G00544521 as “ holotype ”, whereas G00544571 was annotated by him as “ isotype ”. According to the Art. 9.10 of ICN, Barkoudah’s indication of the holotype (which precedes Madhani et al. ’s proposal) and his annotation on G00544521, are errors to be corrected to lectotype. So, G00544521 is the lectotype and P01903171 is the isolectotype for G. frankenioides . Both these specimens matches the protologue and corresponds to the current concept of the species by showing prostrate stems, linear leaves with glandular hairy, oblanceolate pink petals, and inflorescence in apex and sides (YıldızbaŞ 2017).