Bolanthus frankenioides var. fasciculatus (Boiss. & Heldr.) Barkoudah, Wentia

Yildizbaş, Avni & Koç, Murat, 2025, The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye, Phytotaxa 696 (1), pp. 28-54 : 41-42

Bolanthus frankenioides var. fasciculatus (Boiss. & Heldr.) Barkoudah, Wentia 9: 170 (1962) ≡ Gypsophila frankenioides Boiss. var. fasciculata Boiss. & Heldr., Diagn. Ser. 1(8): 58 (1849)

Lectotype [designated by Barkoudah (1962: 170) as “ isotype ”, here corrected according to the Art. 9.10 of ICN): —[ Turkey C3 Isparta] in pascuis siccis montis Stavros Pisidiae supra Isbarta , 10 Aug 1845, Heldreich 1017 ( G00544524 , photo!; ısotype FI, n.v.).

= Bolanthus stenopetalus Hartvig & Strid, In Bot. Jahrb. Syst. 108: 318 (1987)!, syn. nov.

Type: — [ Turkey C2 Muğla]. Sandras Dağ, W. Side of the summit area, 2100–2200 m, snowbed meadows and rocky slopes, serpentine, lat. 37˚04’ N, long. 28˚50’ E, 7 July 1984, Hartvig, Seçmen & Strid 23375 (holotype C; isotypes B, E! EGE, G00226570 !, photo!) .

Typification of the name Gypsophila frakenioides var. fasciculata :— Barkoudah (1962: 170) stated “Type: Turkey: Wilayet Antalya, Mt. Davros-dagh, Heldreich s. n. 1895, isotype ( FI)”. However, no holotype or isotype was reported in the protologue. According to the Art. 9.10 of ICN, Barkoudah’s indication of the isotype is an error to be corrected to lectotype. So, G 00544524 is the lectotype and specimen at FI (see Madhani et al. 2024: 1245) isolectotype for G. frakenioides var. fasciculata .

Phenology: June–July, 1550–2200 m, usually around snow beds or flat landed areas.

General Distribution: Endemic. East Mediterranean Element.

Distribution for Türkiye: The Mediterranean region. ( Figs. 8–9 ).

Notes:— Gypsophila frankenioides var. fasciculata was firstly described by Boissier and Heldrich (in Boissier 1849: 58) and later transferred to the genus Bolanthus by Barkoudah (1962). The taxon exhibits quite a few similarity with the Bolanthus frankenioides . The taxon differes from the other taxa in terms of its pedicel length and trichome density. Due to their overlapping distribution areas, these features were determined as varying morphological characters and thus it was deemed suitable to consider this taxon as variety. Also it has been recommended to be placed in the VU B1a category ( IUCN 2017). Its distribution in Türkiye was given in Figure 9 .

Bolanthus stenopetalus was firstly described by Hartvig & Strid (1987: 318). The location of the type of this taxon is The Sandras Mountain of Muğla Province and at this location, this type is known to be found in the taxon Bolanthus frankenioides var. fasciculatus ( Huber-Morath 1967) . The type specimen of the taxon and the specimens collected during the field studies at Sandras mountain of Muğla province were detaily examined. The paper published by Hartvig and Strid and the detailed descriptions were gathered and as a result, the features that were defined as distinctive characters including ciliated and long leaves, flowers with short pedicels, pink and reverse lanceolate petals and ovariums with 8-10 ovules were identified as the features of the taxon Bolanthus frankenioides var. fasciculatus . For this reason, the taxon was considered as synonym.

Specimens examined: — TÜRKİYE. C2 Muğla Köyceğiz, Sandras mountain, BeŞparmak hill rocky stony soil, 1900 m, 12 July 1978, Özhatay & Özhatay (ISTO-32912) ; Muğla Köyceğiz, eastern slopes of Çiçekbaba hill, 1900 m, 14 July 1978, Özthatay & Özhatay (ISTO-20482) ; Muğla: Sandras mountain, Çiçekbaba hill, serpentine cliff, 2200 m, 26 July 1977, Özhatay No: 1312 (ISTO-19455) ; Muğla: Sandras mountain, BeŞparmak hill, 1900 m, 12 July 1978, Özhatay & Özhatay (ISTE-40513) ; Muğla / Antalya: Akdağ , pebbled slopes, limestone, 1700–2000 m, 08 July 1992, Gemici, Özel & Görk (EGE-26644) ; Muğla: Sandras mountain, SW of the summit, Marshy places by a creek in opening of Pinus nigra forest, serpentine, 1750m, 06 July 1984, Hartvig, Seçmen & Strid (EGE-27895) ; Antalya: Elmalı, Akdağ , 14 July 1968, Pamukçuoğlu (HUB-03415) ; Muğla: Köyceğiz, Yayla village, top of Gökçeova lake , Sandras mountain , serpentine rocky, 2030 m, 05 August 2015, Koç 2188 & Hamzaoğlu (Bozok Univ. Herb.) ; C4 Konya: SeydiŞehir, south of Tınaz mountain , the open of SaydaŞ forest, 1550 m, 20 June 1982, Ocakverdi (KNYA-1216) .

Figures

FIGURE 8. Bolanthus frankenioides var. fasciculatus (Koç 2188 & Hamzaoğlu). A—habit, B—inflorescence, C—flower, D—petal, E—capsule.

FIGURE 9. Distribution of taxa Bolanthus thymoides (D), B. frankenioides var. frankenioides (▲)and B. frankenioides var. fasciculatus (●) in Türkiye.