Drymaria cordata (L.) Willd.

Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas & Montesinos-Tubée, Daniel B., 2025, A nomenclator of Drymaria (Caryophyllaceae), PhytoKeys 266, pp. 139-175 : 139-175

10 a. Drymaria cordata (L.) Willd. in Roem. & Schult., Syst. Veg. 5: 406. 1819. subsp. cordata

≡ Holosteum cordatum L. , Sp. Pl. 1: 88. 1753. Type: Tab. 11 “ Alsine americana nummulariae folio ” in Hermann, Paradisus Batavus, 1698 ( lectotype, designated by Burger in Cafferty and Jarvis 2004, pg. 1052).

= Holosteum diandrum Sw. , Prod. 27. 1788. Type: Jamaica. s. d., O. Swartz s. n. ( holotype: S [ S 10-22103 image!]; isotypes: G [ G 00215840 image!], S [ S-R-2936 image!]) .

≡ Drymaria diandra (Sw.) Macfad. , Fl. Jamaica 1: 52. 1837. nom. illeg. [non D. diandra Blume (1825) ].

≡ Drymaria cordata var. diandra (Sw.) Griseb. , Fl. Brit. W. I. 56. 1859.

= Drymaria cordata var. puberula Triana & Planch. , Ann. Sci. Nat., Bot., sér. 4, 17: 148. 1862. Type: Colombia. [ Cundinamarca]: Bogota , alt. 2600 m, s. d., J. J. Triana s. n. ( lectotype, designated by Duke 1961, pg. 251: P n. v.; isolectotype NY [3578908 image!]) .

= Drymaria procumbens Rose , Contr. U. S. Natl. Herb. 1 (9): 304. 1895. Type: Mexico. Colima: near water ditches about Colima , 9 Jan. – 6 Feb. 1891, E. Palmer 1165 ( holotype: US [15185/00103359 image!]; isotypes: K [ K 000471743 image!], NY [00342506 image!], US [1391060 / 00955563 image!]) .

= Drymaria adenophora Urb. , Repert. Spec. Nov. Regni Veg. 21 (8–20): 213. 1925. Type: Cuba. Pinar del Río: Cortez , 27 March 1921, L. E. Ekman 12798 ( lectotype, designated here: S [ S-R-1684 image!]; isolectotypes: G [ G 00226766 image!], NY [00073724 image!]) .

≡ Stellaria adenophora (Urb.) Leon , Contr. Ocas. Mus. Hist. Nat. Colegio “ De La Salle ” 9: 4. 1950.

= Drymaria sessilifolia Fiori , Anal Bot. Fieni Imp.: 20. 1939. Type: Ethiopia. Adis Abeba, Olettà , alt. 1500 m, 19 October 1937, E. Bartolozzi 13 ( holotype: FT [ FT 001060 image!]) .

= Drymaria cordata var. pacifica M. Mizush. , J. Jap. Bot. 32 (3): 78. 1957. Type: Ecuador. Galapagos Islands: Academy Bay , Indefatigable Islands, 100 ft [alt. 30.48 m], 1 April 1930, H. K. Svenson 65 ( holotype: GH [00037749 image!]) .

Distribution.

Subspecies of wide distribution, from the United States to South America and the Caribbean.

Notes.

Mizushima (1957) chose the specimen at Herb. Linn. no. 109-1 as the lectotype of Drymaria cordata . However, this sheet, as mentioned by Burger in Cafferty and Jarvis (2004), is a post- 1753 addition to the Linnaean herbarium and consequently cannot be considered original material and is ineligible for typification of the name.

Although the illustration designated as type is not of high quality in showing some details of the flowers, e. g., the petals appear undivided, there is no doubt that it corresponds to the concept of Drymaria cordata subsp. cordata as currently recognized.

There are two sheets of Drymaria diandra at S. The specimen considered here as an isotype is apparently a fragment of the holotype that was later removed from the specimen.

Regarding the type of Drymaria cordata var. puberula, Duke (1961) used the term holotype for the Triana specimen at P cited above; however, various syntypes were mentioned in the protologue without a formal designation of type. Thus, Duke’s statement should be treated as an error to be corrected to a lectotype.

Most of the type specimen deposited at B were destroyed in World War II ( Hiepko 1987). Because the holotype in B of Drymaria adenophora was destroyed, we here designated the specimen at S as the lectotype.

For the same reason as with Drymaria arenarioides , authorship of the Drymaria cordata combination should be attributed to Willdenow (see Turland et al. (2018), art. 46.3 example 15).