Tipula (Vestiplex) balioptera Loew, 1863

Starkevich, Pavel, Salmela, Jukka, Olsen, Kjell Magne & Andersson, Michael, 2025, New synonyms of Tipula (Vestiplex) balioptera Loew, 1863 (Diptera, Tipulidae), Biodiversity Data Journal 13, pp. e 141363-e 141363 : e141363-

Tipula (Vestiplex) balioptera Loew, 1863

Tipula balioptera Loew, 1863 - Loew 1863: 284; Alexander 1915: 460;

Tipula (Vestiplex) balioptera : Crampton et al. 1942, Alexander 1965, Starkevich and Paramonov 2016, Starkevich et al. 2020, Brodo et al. 2023, Kolcsár et al. 2023;

= Tipula (Vestiplex) laccata laccata : Savchenko 1964 nec Lundstrom and Frey 1916 (partim), after Starkevich and Paramonov (2016);

= Tipula (Vestiplex) tchukchi : Mannheims 1971 nec Alexander 1934 (partim), after Starkevich and Paramonov (2016). According to Mannheims (1971), T. (V.) tchukchi = T. (V.) laccata : Lackschewitz 1936 (partim); Mannheims 1953 (partim); Savchenko 1964 (partim); nec Lundstrom and Frey 1916;

= Tipula (Vestiplex) tchukchi : Salmela 2008, Salmela 2011, Salmela 2012, Salmela and Petrašiūnas 2014;

Tipula (Vestiplex) bo Mannheims, 1967 - Mannheims 1967: 148 syn. nov. stat. res. Synonymy with T. (V.) tchukchi Alexander, 1934 after Mannheims and Savchenko (1973) = laccata Lackschewitz 1936 (partim), Mannheims 1953 (partim), Savchenko 1964 (partim), nec Lundstrom and Frey 1916;

Tipula perretti Alexander, 1928 - Alexander 1928: 98 syn. nov.; Tipula (Vestiplex) perretti : Crampton et al. 1942.

Materials

Type status: Lectotype. Occurrence: recordedBy: Kennicot; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Canada; verbatimLocality: English River; Record Level: institutionCode: MCZ

Type status: Other material. Occurrence: recordedBy: I. Kakko; individualCount: 2; sex: male, female; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Bombovarri SE; Event: verbatimEventDate: 15 VII 2010; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU391

Type status: Other material. Occurrence: recordedBy: I. Kakko; individualCount: 2; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Bombovarri SE; Event: verbatimEventDate: 15 VII 2010; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU392

Type status: Other material. Occurrence: recordedBy: A. Haarto; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Urtasvaara; Event: verbatimEventDate: 15 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU408

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Tomuttioja; Event: verbatimEventDate: 11 VI – 19 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU824

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: female; associatedSequences: BOLD sample ID JES- 20110086; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Pahtavaara; Event: verbatimEventDate: 11 VI – 19 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU1858

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; associatedSequences: BOLD sample ID JES- 20110129; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Tomuttioja; Event: verbatimEventDate: 11 VI – 19 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU1896

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Utsjoki, Galddasjohka; Event: verbatimEventDate: 18 VI – 19 VII 2007; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU825

Type status: Other material. Occurrence: recordedBy: A. Rinne; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Utsjoki, Teno; Event: verbatimEventDate: 22 VI – 3 VII 2006; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU851

Type status: Other material. Occurrence: recordedBy: A. Järvinen; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Troms, Storfjord, Skibotn, Veajetnibba S; Event: verbatimEventDate: 24 VI – 16 VII 2017; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.ins2018-168

Type status: Other material. Occurrence: recordedBy: A. Järvinen; individualCount: 1; sex: male; occurrenceID: 8016767E-E022-557C-BF0F-F718132F850F; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Veajetnibba S; Event: verbatimEventDate: 19 VII – 12 VIII 2017; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.ins2018-199

Type status: Other material. Occurrence: recordedBy: A. Järvinen; individualCount: 3; sex: 2 males, 1 female; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Veajetnibba S; Event: verbatimEventDate: 16 VII – 19 VII 2017; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.ins2018-205

Type status: Other material. Occurrence: catalogNumber: 721756 ; recordedBy: K. M. Olsen; individualCount: 1; sex: male; occurrenceID: F7C421F7-3612-5A86-8672-1B18AE8B97C9; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Troms, Kåfjord, N Nuorrtit Gussačohkka; Event: verbatimEventDate: 12 VII 2022; Record Level: institutionCode: PCKMO

Type status: Other material. Occurrence: catalogNumber: 718592 ; recordedBy: K. M. Olsen; individualCount: 5; sex: 3 males, 2 females; occurrenceID: 6797498A-FFE2-5196-A7AB-00AA10C38760; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: N Nuorrtit Gussačohkka; Event: verbatimEventDate: 1 VII – 12 VII 2022; Record Level: institutionCode: PCKMO

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18002 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 3 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18001 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 9 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18004 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 9 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18006 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: female; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 9 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18003 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 15 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18005 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Bojobæski; Event: verbatimEventDate: 16 VII 1924; Record Level: institutionCode: TROM

Type status: Holotype. Occurrence: catalogNumber: No. 2559 ; individualCount: 1; sex: male; preparations: Antenna and wing on slide mounted, USNM; Taxon: scientificName: Tipula (Vestiplex) perretti Alexander, 1928 ; Location: country: Canada; verbatimLocality: Hopedale, Labr; Record Level: institutionCode: CNC

Type status: Paratype. Occurrence: catalogNumber: Type # 6365: 2; MZLU- 110421 ; recordedBy: A & S Ulfstrand; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) bo Mannheims, 1967; Location: country: Sweden; verbatimLocality: Lycksele Lappmark, Vindelälven, N Tjatsemvare, 45 km NW Ammarnäs; Event: verbatimEventDate: 23 VII 1965; Record Level: institutionCode: MZLU; source: https://www.flickr.com/photos/tags/mzlutype06365/

Type status: Paratype. Occurrence: catalogNumber: Type # 6365: 3; MZLU- 110422 ; recordedBy: A & S Ulfstrand; individualCount: 1; sex: male; Taxon: scientificName: Tipula (Vestiplex) bo Mannheims, 1967; Location: country: Sweden; verbatimLocality: Lycksele Lappmark, Vindelälven, N Tjatsemvare, 45 km NW Ammarnäs; Event: verbatimEventDate: 23 VII 1965; Record Level: institutionCode: MZLU; source: https://www.flickr.com/photos/tags/mzlutype06365/

Type status: Paratype. Occurrence: catalogNumber: Type # 6365: 4; MZLU- 110423 ; recordedBy: A & S Ulfstrand; individualCount: 1; sex: female; Taxon: scientificName: Tipula (Vestiplex) bo Mannheims, 1967; Location: country: Sweden; verbatimLocality: Lycksele Lappmark, Vindelälven, N Tjatsemvare, 45 km NW Ammarnäs; Event: verbatimEventDate: 23 VII 1965; Record Level: institutionCode: MZLU; source: https://www.flickr.com/photos/tags/mzlutype06365/

Taxon discussion

The first record of T. (V.) balioptera out of the Nearctic Region was published by Starkevich and Paramonov (2016) with taxonomic account as follows:

= Tipula (Vestiplex) laccata laccata : Savchenko 1964 nec Lundstrom and Frey 1916 (partim), after Starkevich and Paramonov (2016).

A single male specimen from Yakutyia, Russia was detected amongst material identified as T. (V.) laccata by E. N. Savchenko ( Savchenko 1964). Another partim of the T. (V.) laccata material belonged to T. (V.) aldrichiana , a species which was also recorded in the Palaearctic for the first time ( Starkevich and Paramonov 2016). Later, T. (V.) balioptera was recorded in Mongolia ( Starkevich et al. 2020) and Norway ( Kolcsár et al. 2023).

Following Mannheims (1971), all material published by Savchenko (1964) as T. (V.) laccata was synonymised with T. (V.) tchukchi Alexander, 1934 , including some other material listed below:

Tipula (Vestiplex) tchukchi Alexander, 1934

= laccata Lackschewitz 1936 (partim), Mannheims 1953 (partim), Savchenko 1964, nec Lundstrom and Frey 1916, after Mannheims (1971).

Finally, another species T. ( V.) bo Mannheims, 1967 was synonymised by Mannheims and Savchenko (1973) with the taxonomic list as follows:

Tipula (Vestiplex) tchukchi Alexander, 1934

= laccata Lackschewitz 1936 , Mannheims 1953, Savchenko 1964, nec Lundstrom and Frey 1916, after Mannheims and Savchenko (1973).

As can be seen from the taxonomic records ( Mannheims 1971, Mannheims and Savchenko 1973), besides T. ( V.) bo, three partims of T. (V.) laccata material were listed as T. (V.) tchukchi .

After examination of type material of T. ( V.) bo and based on morphology analysis, we found that this species belongs to T. (V.) balioptera and thus is a junior synonym:

Tipula (Vestiplex) bo Mannheims, 1967 - Mannheims 1967: 148 syn. nov. stat. res.

= laccata Lackschewitz 1936 (partim), Mannheims 1953 (partim), Savchenko 1964 (partim), nec Lundstrom and Frey 1916.

Subsequently, T. (V.) tchukchi is exluded from the list of fauna from Norway, Sweden and Finland.