Speophilus kiesenwetteri castellsaperai Jeannel, 1924

Prieto-Manzanares, Miguel & Masó-Ros, Glòria, 2025, Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain), Zootaxa 5682 (1), pp. 1-102 : 80

Speophilus kiesenwetteri castellsaperai Jeannel, 1924 ( Figs. 21 , 65 )

Speophilus Kiesenwetteri subraza Castellsaperai Zariquiey, 1924: 10; fig. 4. (unavailable)

Speophilus (Speophilus) Kiesenwetteri subsp. Castellsaperai Jeannel, 1924a: 171.

Current accepted name

Troglocharinus sanllorensi ( Zariquiey, 1924) : Fresneda et al., 2024: 776.

Type locality

Avenc de Castellsapera [original citation: “Avenc de Castellsapera, Sant Llorenç de Munt”].

Type material

Syntypes: MZB 78-1194, MZB 78-1455 / MZB 78-1461, MZB 78-2708, MZB 78-2709 (5 ♂♂, 5 ♀♀), type loc., 25-XI-1922, S. Novellas leg. (ex coll. R. Zariquiey).

Remarks

Zariquiey (1922a) in a previous publication announced the discovery of a new taxon (a subspecies of Speophilus kiesenwetteri ), based on four specimens collected by A. Codina on 12 October 1913, but provided no description. Subsequently, Zariquiey (1924) proposed the name castellsaperai as a infrasubspecific entity (“subraza”) as is shown in the publication (article 45.6.1.) in which he also described another new entities explicitly as subspecies of Speophilus kiesenwetteri . The name castellsaperai was used at a later date by Jeannel (1924a), which made it available for a subspecies of S. kiesenwetteri, Jeannel thereby becoming the author of this new subspecies (article 45.5.1.).

Zariquiey (1924) mentions “bastantes ejemplares” [plenty of specimens] collected by S. Novellas, of which we have located 10 in the MCNB collection, bearing red labels printed with “ Typus ” (the only exception is one labelled as “COTIPO”). Zariquiey also cites an indeterminate number of specimens collected by Faura i Sans and the four above-mentioned specimens collected by Codina. The specimens of the type series collected by Faura i Sans were deposited in the Biospeologica collection ( Zariquiey, 1924; Jeannel, 1924a) that is currently in the MNHNP. In fact, in box 73 of the general collection of the MNHNP we have found two of these specimens, labelled as cotypes, as well as one of the specimens of the series captured by Novelles (with identical capture labels to those of the specimens deposited in the MCNB and with the same red printed labelled indicating “ Typus ”). After Zariquiey’s description (1924), Codina (1924) cites three “Tipus” along with nine other specimens from the type locality with a capture date of 12 October 1913 deposited in his private collection. However, we have failed to find in the MCNB collection any of the specimens captured by Codina. In the MNCNM collection (box 27), we have found a specimen with an identical capture label to those from the series in the MCNB and with a red label printed with the inscription “Cotypus”.

Figures

FIGURES 20–22. Original labels associated to type specimens of Iberian Leptodirini: 20A–B. Speophilus jacasi Lagar, 1966, Cova de les Rondes: 20A. paratype ♀ (MZB 78-1215); 20B. paratype (MZB 78-1241); 21. Speophilus kiesenwetteri castellsaperai Jeannel, 1924, syntype ♀, Avenc de Castellsapera (MZB 78-1194); 22A–B. Troglocharinus españoli españoli Zariquiey, 1950, Avenc d’Ancosa: 22A. syntype ♂ (MZB 78-2041); 22B. syntype ♀ (MZB 78-2919).

FIGURES 62–67. Type specimens of Iberian Leptodirini. Habitus photographs, dorsal view (scale bar: 1 mm): 62. Speophilus cenarroi Español,1955, holotype ♂,Forat de les Gralles (MZB 78-2067); 63. Speophilus españoli Jeannel, 1930, syntype♀, Cova del Traça (MZB 78-1214); 64. Speophilus jacasi Lagar, 1966, paratype ♀, Cova de les Rondes (MZB 78-1215); 65. Speophilus kiesenwetteri castellsaperai Jeannel, 1924, syntype ♀, Avenc de Castellsapera (MZB 78-1194); 66. Speophilus subilsi Español, 1966, syntype ♂, Avenc de l’Obaga del Grau (MZB 78-1317); 67. Speophilus kiesenwetteri sanllorensi Zariquiey, 1924, syntype ♂, Cova de Santa Agnès (MZB 78-1462).