Isapis pulcherrima (Herrich-Schäffer, [1853]), Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomic analysis reveals new species and subspecies of butterflies, The Taxonomic Report of the International Lepidoptera Survey 11 (6), pp. 1-63 : 23

Eurygona? pulcherrima Herrich-Schäffer, [1853] and Themone poecila H. Bates, 1868 belong to the genus Isapis E. Doubleday, 1847 and not Themone Westwood, 1851

Eurygona? pulcherrima Herrich-Schäffer, [1853] (type species in Suriname) and Themone poecila H. Bates, 1868 (type locality Brazil: Amazonas, Ega [= Tefé]) currently placed in the genus Themone Westwood, 1851 (type species Helicopis pais Hübner, [1820] ) are not monophyletic with it and instead form a clade together with Isapis E. Doubleday, 1847 (type species Papilio agyrtus Cramer, 1777 ) and are close to it genetically ( Fig. 21). To restore monophyly of Themone , we transfer the two species to Isapis forming new combinations Isapis pulcherrima (Herrich-Schäffer, [1853]) , comb. nov. and Isapis poecila (H. Bates, 1868) , comb. nov. Furthermore, we note that the clade with these species is sister to Melanis Hübner, [1819] (type species Papilio melander Stoll, 1780 ), and they are closely related to it, i.e., COI barcodes of the type species of Melanis and Isapis differ by 7.3% (48 bp). Therefore, Isapis can be included in Melanis , a step we are not taking here but proposing for consideration.