Hymenopus heteromorphus (Benth.) Sothers & Prance var. heteromorphus, Kew Bull.

Barros, Natália, Menezes, Luis Fernando Tavares de, Sothers, Cynthia & Peixoto, Ariane Luna, 2025, Flora of Espírito Santo: Chrysobalanaceae, Rodriguesia (e 00872024) 76, pp. 1-39 : 23

5.1. Hymenopus heteromorphus (Benth.) Sothers & Prance var. heteromorphus, Kew Bull. 71(58): 18. 2016.

Basionym: Licania heteromorpha Benth., J. Bot. (Hooker) , 2: 221. 1840. Figs. 1 g-h; 2

Tree, up to 26 m tall, DBH 18–55 cm., stem ca. 5–16 m tall; young branches hispid, becoming glabrous with age, lenticellate. Leaves elliptic to oblong-elliptic, 7.7–10.2 × 3.2–4.7 cm, chartaceous to coriaceous, slightly discolorous, base cuneate, with 2 glands, apex rounded to acute, glabrous on both sides, midrib prominulous adaxially, sparsely hispid on both surfaces, becoming glabrous with age adaxially, without stomatal crypts, margin glabrous to hispid, becoming glabrous with age, primary veins 10–13 pairs, sparsely hispid abaxially; petiole 4–6 mm long, hispid, terete; stipules 2, lanceolate, ca. 5.5 mm long, chartaceous, hispid abaxially, caducous. Inflorescence subterminal to terminal, panicle of cymules, pubescent; bracts caducous (not seen); bracteoles 1–1.5 mm long, triangular, pubescent. Flowers 2–5 mm long, white, green or yellow, pubescent, sessile or pedicellate, pedicel 1–1.5 mm long; receptacle campanulate, 1–1.5 mm long, pubescent; sepals triangular, ca. 1 mm long, glabrous adaxially, pubescent abaxially;petals 5, ca. 1 mm long, glabrous adaxially, pubescent abaxially; stamens 5, filaments ca. 0.5 mm long, glabrous, free, included, inserted in a complete circle; ovary ca. 0.5 mm long, inserted at receptacle base, villous; style ca. 0.5 mm long, glabrous. Drupe globose, sessile, 2–2.7 × 2.2–2.5 cm, epicarp granular, glabrous.

Selected material examined: Conceição da Barra , Aracruz Celulose S.A., 22.IX.1992,fr., O.J.Pereira 3883 ( VIES) ; Itaúnas , mata seca de restinga sobre cordão arenoso próximo de uma área entre cordões dominado por Montrichardia , área atrás da Fazenda Jequitaia, 14.I.2010, fl., A. Giaretta 692 ( SAMES) . Linhares, Reserva Natural Vale , 14.XII.1981, fl., H.C. de Lima 1660 ( NY, RB) . Jaguaré, lado esquerdo da casa de guarda na área do Estivado , 30.XI.1992,fr., D.A. Folli 1744 ( RB) . Marilândia, Alto Liberdade,propriedade de Deuclecio Lorenzini. 19.IV.2006, fr., L.F.S. Magnago 943 ( MBML) . Pinheiros, Rebio Córrego do Veado , 13.II.2022, L.F.T Menezes 2937 ( SAMES) . Santa Teresa, Rebio Augusto Ruschi, Nova Lombardia , 7.XI.2001, fr., L. Kollmann 4985 ( MBML) .

Hymenopus heteromorphus var. heteromorphus is quite different from the other taxa occurring in the state. The flowers and its structures are reduced, especially the included stamens. In some exsiccatae there are pedicellate and sessile flowers in the same inflorescence. The slightly discolorous leaves and the hispid pubescence present on young branches, stipules, leaf margin and primary veins also help in its identification. Collected with flowers from March to July and December, and fruits from April and September to November.

This taxon occurs in Bolivia, Brazil, Colombia, French Guiana, Guyana, Peru and Venezuela ( Sothers et al. 2016). In Brazil it is recorded in Amazonas, Amapá, EspÍrito Santo, Maranhão, Mato Grosso, Pará, Rio de Janeiro, Rondônia and Roraima ( Barros et al. 2022; Sothers & Prance 2023d). In EspÍrito Santo it occurs in Restinga, Ombrophilous forest and Mata de Tabuleiro, in Conceição da Barra, Linhares,Jaguaré, Marilândia, Pinheiros and Santa Teresa municipalities. In the state it was collected in three protected areas (Reserva Natural Vale, Reserva Biológica Córrego do Veado, Reserva Biológica Augusto Ruschi) and can be considered “Near Threatened” ( NT).

The common name for this species is Carrapeta (D.A. Folli 1744, D.A. Folli 4801, I.A. Silva 008).