JURELLANIDAE KLOMPMAKER & ROBINS, 2019

Robins, Cristina M. & Klompmaker, Adiël A., 2019, Extreme diversity and parasitism of Late Jurassic squat lobsters (Decapoda: Galatheoidea) and the oldest records of porcellanids and galatheids, Zoological Journal of the Linnean Society 187, pp. 1131-1154 : 1136

FAMILY JURELLANIDAE KLOMPMAKER & ROBINS FAM. NOV.

urn:lsid:zoobank.org:act:EE13BE21-7CE1-4A1B-82E3-83EF9097B01D

Type genus: Jurellana Schweitzer & Feldmann, 2010b .

Other included genera: Ovalopus Klompmaker & Robins , gen. nov.

Diagnosis: Ovate carapace, flattened for largest specimens, widest near mid-length, equally long as wide to ~10% longer than wide (excluding rostrum); rostrum not sulcate axially; fronto-orbital width ~70–80% of maximal width; orbital cavity with vertical ridge, anterolaterally oriented, weak outer-orbital spine directed forward; lateral rim present anteriorly; protogastric and hepatic regions confluent; grooves weak to moderately strong; cervical and branchiocardiac grooves parallel or nearly parallel; post-cervical groove present.

D i s c u s s i o n: J u r e l l a n i d a e f i t s b e s t i n Homolodromioidea because of a combination of the following characters: the carapace is as long as wide or somewhat longer than wide; the orbital cavity contains a vertical ridge and is anterolaterally oriented; the cervical and branchiocardiac grooves continue onto the flank; presence of a groove curving forward around the subhepatic region; and the presence of a post-cervical groove. For Jurellana , a merging of the branchiocardiac groove with the cervical groove has not been confirmed, and the subhepatic region is not swollen, both of which are mentioned to be generally the case in homolodromioids (see diagnosis by Karasawa et al., 2011: 534–535). However, the presence of an inflated subhepatic region has not been confirmed for many species of the homolodromioid Tanidromites Schweitzer & Feldmann, 2008a , and the branchiocardiac groove of Tanidromites and Eodromites Patrulius, 1959 can be interrupted on the flank ( Schweitzer & Feldmann, 2008a: pls. 4B, 6D, 6F; A.A.K., pers. obs.).

Jurellanidae differs from all other homolodromioid families. Bucculentidae have swollen hepatic regions representing the widest parts of the dorsal carapace and have well-delineated regions. The variable Goniodromitidae bear rostra that are often axially sulcate, have generally steeper flanks, and appear less flattened for equally sized specimens. Homolodromiidae have a rostrum with two forwardly directed lateral spines and the carapace widens posteriorly. Longodromitidae generally have a higher length-to-width ratio, the rostrum is axially sulcate, and a clear lateral rim is often absent. Prosopidae widen posteriorly and have well-defined regions. Tanidromitidae exhibit a higher length-to-width ratio and exhibit longer flanks.

GENUS OVALOPUS KLOMPMAKER & ROBINS GEN.