Camptomorpha papillosa ( Attems, 1931 )

Golovatch, Sergei I. & Bouzan, Rodrigo S., 2025, Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae), Arthropoda Selecta 34 (1), pp. 1-15 : 7

Camptomorpha papillosa ( Attems, 1931)

Fig. 7 .

Leptodesmus (Leptodesmus) pApillOsus Attems, 1931: 10 , figs 1–4 (sYntYpes, ZMH); Attems, 1938: 13, fig. 8; Weidner [1960: 84], listed in type catalogue.

CAmptOmOrpHA pApillOsus (sic!) — Hoffman, 1953: 123.

Leptodesmus ( PHAntAsmOdesmus ) pApillOsus — Kraus, 1956: 142, Camptomorpha by implication.

Pseudoleptodesmus (Pseudoleptodesmus) cOrdillerAnus Attems, 1931: 28 , figs 40–42, syn.n. (sYntYpes, ZMH, after Hoffman [in litt.]).

Leptodesmus (Pseudoleptodesmus) cOrdillerAnus — Attems, 1938: 39, fig. 42.

CAmptOmOrpHA cOrdillerAnus (sic!) — Hoffman, 1953: 122.

Leptodesmus ( PHAntAsmOdesmus ) cOrdillerAnus — Kraus, 1956: 142, Camptomorpha by implication.

BRIEF DESCRIPTIVE NOTES (after Attems [1931, 1938]). Width 4.0 mm. Coloration dark castaneous brown with a vague, lighter, axial stripe; antennae and legs dark red-brown. Paraterga relatively small and faintly rimmed, caudal corners mostly sharp and dentiform ( Fig. 7A ). Epiproct simple and finger-shaped, directed slightlY dorsad. HYpoproct subtrapeziform. with a small apical tooth between both thick setigerous knobs. ♂ legs ventrallY with all tibiae but four last pairs carrYing apical chelae; ♂ coxae 3 each with a strong, blunt, denselY setose process directed forward.

Gonopods ( Fig. 7B–F ) with both prefemoral process (PfP) and acropodite (A) subequal in length, very complex and strongly armed; PfP also broad, with an irregular internal margin and a small digitiform process near middle; A robust and leaf-shaped, with several grooves at external margin, one of the apical branches being directed basally; solenomere (sl) simple and only slightly curved.

DISTRIBUTION. Ecuador, Sabanilla, 1°50′45.0″S 80°13′ 04.3″W, East slope of the Cordilleras [ Attems, 1931, 1938].

JUSTIFICATION OF SYNONYMY. Hoffman [in litt.] provided ample evidence of the synonymy of C. pApillOsA with C. cOrdillerAnA ( Fig. 7B–F ). Indeed, both these formal species came from the same collection and even the same population, even though Attems [1931] assigned them to different genera! Following Hoffman [in litt.], we choose to use the name C. papillOsA because it has 26 pages of priority over C. cOrdillerAnA, while both names have been used the same number of times in the primary taxonomic literature.

Figures

Fig. 7. CAmptOmOrpHA pApillOsA (Attems, 1931), ♂ sYntYpes of C. cOrdillerAnA (A–C) and C. pApillOsA (D–F). A — left halves of rings 10 and 11, dorsal view; B — right gonopod, mesal view; C–F — left gonopod, lateral, mesal, sublateral and dorsal views, respectively. Designations: A — acropodite; PfP — prefemoral process; sl — solenomere.After Attems [1931] (A–C) and Hoffman [in litt.] (D–F). Рис. 7. CAmptOmOrpHA pApillOsA (Attems, 1931), синтипы ♂♂ C. cOrdillerAnA (A–C) и C. pApillOsA (D–F). A — левые половины туло- виЩных сегментов 10 и 11, сверху; B — правый гонопод, иЗнутри; C–F — левый гонопод, соответственно сбоку, иЗнутри, пости сбоку и сверху. ОбоЗначениЯ:A — акроподит; PfP — префеморальный отросток; sl — соленомер. По:Attems [1931] (A–C) и Hoffman [in litt.] (D–F).