Tsivoka Villiers, 1982, 2011

Jeong, Soo-Hyun, Vives, Eduard & Mckenna, Duane D., 2025, A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species, Zootaxa 5632 (1), pp. 159-170 : 161-162

The genus Tsivoka Villiers, 1982

Family Cerambycidae Subfamily Cerambycinae Tribe Dorcasomini Subtribe Apatophyseina

Genus Tsivoka Villiers, 1982: 132 , 157, 159

Tsivoka Villiers, Quentin & Vives, 2011: 39 , 342

Tsivoka Švácha & Lawrence, 2014: 134 , 142 Type-species: Tsivoka testaceipes ( Fairmaire, 1889)

Redescription of Tsivoka Villiers, 1892 (Revised from Villiers et al. 2011): Head with an elongate and subparallel rostrum. Genae extended beyond the lower lobe of the eyes, which are finely faceted, narrow and weakly indented. Frons almost flat, divided by a dorsal furrow that extends from the inter-antennary area up to the level of the posterior edge of the eyes, short and subparallel temples. Mandibles large, falcate, inner margin sinuate. Maxillary and labial palps densely covered with fine setae. Antennae inserted in front of the eyes, closer to them than to the base of the mandibles; base of antennae prominent and close together; scape long, curved at apex; antennomere III slightly shorter than scape, subequal to IV; V and the following ones longer than IV, and projected at the external apical angle from VI to X. Strongly convex pronotum with barely marked disc tubercles, the two medians close together, often obsolete, the two posterior ones further apart and a more evident; anterior margins and posterior margins strongly depressed, finely rimmed; almost straight posterior half of sides, strongly convergent forward, this line break forming an obtuse angle. Scutellum as long as it is wide. Elytra strongly narrowed from anterior to posterior, humeri slightly prominent in males, more strongly so in females; apex truncate. Fusiform femora; straight tibiae, metatibiae widened at apex and, in males, with a small process at the end of the inner edge; protarsomeres strongly dilated in males; metatarsomeres I as long or longer than the metatarsomeres II-V together; onychium strongly dilated and toothed in males. Tergite VII of males subrectangular, longer than wide; sternite VII with a wide but shallow indent; sternite VIII short and deeply indented. Apically lobed tegmen very long, arched, and pubescent. Male genitalia of the same type as in Mastododera .

Remarks: Tsivoka has been observed on the flowers of Mimosa latespinosa Lamarck, 1806 ( Fabaceae ) in bright sunlight ( Villiers et al. 2011).

Key to species of Tsivoka (modified from Villiers et al. 2011)

1. Head, pronotum and scutellum black ( Figs 3 , 4 )............................................................. 2

– Head, pronotum and scutellum orange ( Fig. 1b )........................................ T. peyrierasi Villiers, 1982

2. Elytra black with a distinctive bright orange mark restricted to humeral area ( Figs 1a , 3 , 4 ).......... T. humeralis sp. nov.

– Elytra brown to black, without a distinctive orange mark or with longitudinal orange vittae from humeri to apex of elytra (Figs 2, 7)............................................................................................... 3

3. Elytra light brown to black without clear yellow band; Meso and metatibiae yellow or red ( Fig. 2 )................................................................................................ T. testaceipes ( Fairmaire, 1889)

– Elytra black with a narrow yellow band, clearly defined, extending from the shoulder almost to the apex. Meso and metatibiae black or dark brown. ( Fig. 7 ).................................................... T. simplicicollis ( Gahan, 1890)

Figures

FIGURE 1. Specimens and labels of Tsivoka in MNHN. a, T. humeralis sp. nov. female; b, T. peyrierasi Villiers male.

FIGURE 2. Tsivoka testaceipes (Fairmaire). a, male, dorsal view; b, male, lateral view; c, female, dorsal view; d, female, lateral view.

FIGURE 3. Tsivoka humeralis sp. nov. male, holotype. a, dorsal view; b, lateral view; c, head, dorsal view; d, pronotum, dorsal view.

FIGURE 4. Tsivoka humeralis sp. nov. female. a, dorsal view; b, lateral view

FIGURE 7. Tsivoka simplicicollis (Gahan) a, male, dorsal view; b, female, dorsal view.