Crassiclitellata, Jamieson, 1988

Erséus, Christer, Williams, Bronwyn W., Horn, Kevin M., Halanych, Kenneth M., Santos, Scott R., James, Samuel W., Châtelliers, Michel Creuzé des & Anderson, Frank E., 2020, Phylogenomic analyses reveal a Palaeozoic radiation and support a freshwater origin for clitellate annelids, Zoologica Scripta 49 (5), pp. 614-640 : 628

3.3.4 | Crassiclitellata

Relationships within Crassiclitellata closely match those recovered in Anderson et al. (2017 %, despite the smaller taxonomic sample used here. As seen in that study, the positions of the moniligastrid Drawida sp. and the haplotaxid Pelodrilus sp. vary somewhat across analyses. In most trees, Drawida sp. is sister to a clade comprising all sampled crassiclitellates and Pelodrilus sp. , but Pelodrilus is sister to a subclade comprising the North American taxa Komarekiona and Sparganophilus and Kynotus from Madagascar (Figure 1%. In two trees (25% MARE and 75% TreSpEx-and-BaCoCa-filtered%, Pelodrilus is sister to a clade comprising all crassiclitellates plus Drawida sp. , with Drawida sp. recovered as sister to a large clade comprising all crassiclitellates except Komarekiona , Kynotus and Sparganophilus . In two trees (50% MARE and 75% MARE%, Pelodrilus is sister to the Komarekiona + Kynotus + Sparganophilus clade, with Drawida sister to a clade comprising all other crassiclitellates. The only other relationships within Crassiclitellata that vary across trees involve Eudrilus eugeniae ; in most trees, it is sister to a clade comprising three members of Megascolecoidea ( Acanthodrilus , Dichogaster and Pontodrilus % and two ocnerodrilids ( Kerriona and an unidentified worm from Madagascar %, with a clade comprising Andiorrhinus , Areco and Pontoscolex then being sister to that group. By contrast, Eudrilus is recovered as sister to Andiorrhinus sp. in the 75% MARE tree (Figure S3%.