Pseudolysimachion × sapiehae Holub

Albach, Dirk C., 2025, From just a few to the most type-rich herbarium for Veronica L. (Plantaginaceae) - The effect of digitization of the Muséum national d’Histoire naturelle herbarium in Paris, Adansonia (3) 47 (7), pp. 47-130 : 75

Pseudolysimachion × sapiehae Holub in Holub & Pouzar

Folia Geobotanica et Phytotaxonomica 2: 426 ( Holub & Pouzar 1967). — V. × sapiehae (Holub) Albach , Taxon 57: 5 ( Albach 2008a).

TYPE CITATION. — Ukraine, Ternopil: “Schultz herb. norm. n. ser. Cent. 21, no. 2052 – Veronica incana x spuria Blocki , V. sapiehae Bl. – Julio 1884 – In pascuis gypsaceis ad ripam fluvii Seret prope Olexince [= Oleksyntsi], inter parentes. Galicia. Austria. Leg. Dr. A. Błocki”.

HOLOTYPE. — PR [ PR222519 !].

ISOTYPES (not mentioned byHolub & Pouzar 1967). — BP[ BP 159398 !, BP159399 !, BP 349966 !], M[ M0188531!* ], MPU![2 pls], OXF!, P[ P 04051000 * , P05425258* ].

Nomenclatural note. Błocki used three different names ( V. sapiehae , V. kerneria , V. polonica ) for plants collected in Oleksyntsi (Western Ukraine) and identified as hybrids between V. incana and V. spuria . Since all three names by Błocki are herbarium names, it is not clear, whether he distinguished those by morphological characters or whether he considered them to be synonymous. No obvious morphological difference exists. However, the fact that Błocki used terms “superspurio” and “subspurio” on the labels indicates that he saw more or less influence of V. spuria . The name “sapiehae ” seems to be used only in Schultz’ Herbarium Normale for plants collected by Błocki in july 1884 whereas V. kerneri and V. polonica is on handwritten labels by Błocki with the former used on specimens between 1882 to 1894. However, Holub & Pouzar (1967) only referred to V. sapiehae Błocki when introducing the name Pseudolysimachion x sapiehae Holub. The hybrids cannot be found at the locus classicus anymore. Therefore, the type material is important to verify the hybrid status.