Veronica velenovskyi Uechtr. ex Bornm. var. subintegrifolia Borbás

Albach, Dirk C., 2025, From just a few to the most type-rich herbarium for Veronica L. (Plantaginaceae) - The effect of digitization of the Muséum national d’Histoire naturelle herbarium in Paris, Adansonia (3) 47 (7), pp. 47-130 : 88

Veronica velenovskyi Uechtr. ex Bornm. var. subintegrifolia Borbás

Herbarium Normale Schedae Centuriarum XXXV: 144, nr. 3439 ( Dörfler 1898).

TYPE CITATION. — Hungary, Budapest: “Hungaria centr. In humidis calidis ad ‘Bekás-Megyer’ proper ‘Budapest’, Septembri 1894. Leg. V. v. Borbás et L. Richter”.

LECTOTYPE (designated here). — WU[ WU0152672 !].

ISOLECTOTYPES (designated here). — B[ B 10 0367913! ], BREM!, E!, M![2 sheets], MPU!, OXF!, P[ P03520492* ], PRC[ PRC453965 !, one more sheet!], W[ W0222328! ], Z!

Nomenclatural note. Veronica velenovskyi var. subintegrifolia , the entire-leaved and longer stemmed variety of V. velenovskyi , may be a type growing in submerged water with the serrate type a terrestrial form. Veronica velenovskyi is a synonym of V. scardica , which is a serpentine specialist related to V. anagallis-aquatica . The species is known from scattered localities between Austria and Iran and south to Egypt, although the common origin is unclear and parallel adaptation to serpentine soil is possible, especially of the Egyptian plants ( Ellmouni et al. 2017). Borbás collected his type material near Budapest, whereas the type of V. velenovskyi is from the Bulgarian Black Sea Coast and that of V.scardica from Albania.Strangely, type material of Borbás name was not found in BP. The Herbarium Normale was distributed from Vienna by Dörfler in 1894 and since specimens from other collectors were not part of Dörfler’s personal herbarium ( Vogt et al. 2018), the best specimens are likely to be found in Vienna.