Veronica aleppica Boiss. var. schizostegia Bornm.

Albach, Dirk C., 2025, From just a few to the most type-rich herbarium for Veronica L. (Plantaginaceae) - The effect of digitization of the Muséum national d’Histoire naturelle herbarium in Paris, Adansonia (3) 47 (7), pp. 47-130 : 106

Veronica aleppica Boiss. var. schizostegia Bornm.

Repertorium specierum novarum regni vegetabilis 9: 113 ( Bornmüller 1910). — V. aleppica subsp. schizostegia (Bornm.) Bornm. Beihefte zum botanischen Centralblatt 28: 480 ( Bornmüller 1911). — V. schizostegia (Bornm.) Doostm. & Bordbar Biology 11: 639 ( Doostmohammadi et al. 2022).

TYPE CITATION. — Iraq, prov. Arbil: “Kurdistania Turcica (Assyria): in monte Kuhi-Sefin supra pagum Schaklava, 1000 m. s. m. (6. V. 1893 legi; exsicc. no. 1628). — Kurdistania persica: In monte Pir-Omar-Gudrun (VI. 1867, leg. Haussknecht) et [ Iran, prov. Kerman] inter Kerind et Kermanschah /Gawarreh in monte Kuh-i-Dalahu (9. VI. 1906, leg. Th. Strauss”.

LECTOTYPE. — Designated by Fischer (1981); second-step lectotype designated by Doostmohammadi et al. (2022: 22), B[ B 10 0278579 !].

ISOLECTOTYPES. — Designated by Doostmohammadi et al. (2022: 22), B[ B 10 0278578 !], BP[ BP347767 !], BR[ BR0000005423033 !], JE[ JE00000152 !], OXF!, P[ P03529531 *, P03529532 *], W[ W1895- 1676 !], WU[ WU029659 !].

Nomenclatural note. Veronica schizostegia was initially considered by Bornmüller to be a distinct species, which is the name under which the type material can be found in herbaria, but he changed his mind in his publications and called it either a variety ( Bornmüller 1910) or subspecies of V. aleppica ( Bornmüller 1911) . The taxon was later ( Riek 1935) considered to be a synonym of V. macrostachya . However, phylogenetic analyses of DNA sequence data revealed that these plants from Kurdish parts of Iraq and Iran do not form a monophyletic group with Anatolian and Levantine specimens of V. macrostachya , thus, confirming Bornmüller’s initial thought. Bornmüller (1910) stated three collections of three collectors for his name but only two specimens of his own collection are in P.