Penthicus acuticollis (Reitter, 1887)
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publication ID |
https://doi.org/10.11646/zootaxa.5573.1.1 |
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publication LSID |
lsid:zoobank.org:pub:C2A78D6D-4209-418F-9D90-6ACE3FE9942F |
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persistent identifier |
https://treatment.plazi.org/id/B11AEC76-FFE4-8218-FF0A-26BBFDDD6C7F |
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treatment provided by |
Plazi |
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scientific name |
Penthicus acuticollis |
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acuticollis species-group
Material (studied). P. acuticollis ( Reitter, 1887) ( Figs 5 View FIGURE 5 , 6 View FIGURE 6 ): China, Xinjiang (N Kunlun Mts.): 4 ♂♂ ( ZIN): “Polu 1/VI 90 Gr.” [Kunlun Mts. E Keria, 1.vi.1890, leg. B.L. Grombchevsky]; 1 ♂, 1 ♀ ( ZIN): “Raskem, 30/IX 89 Gr” [ Raskem-Darya River , 30.ix.1889, leg. B.L. Grombchevsky] ; 1 ♀ ( ZIN): “ Ur. Karasay (khr. Russk) V-90 Pevtsov” [ Russian label: natural boundary Karasay (Russky ridge), v.1890, leg. M.V. Pevtsov] .
P. tzaidamicus ( Bogatchev, 1951) ( Fig. 7 View FIGURE 7 ): lectotype, ♂ ( ZIN), designated here: “Tsaidam r. Boming-gola 25– 26.VI.1895 ex. Kozlova” [handwritten; now in China, Qinghai Province, Qaidam Prefecture ], “Coll. Semenov-Tian- Shansky” [print], “ Lobodera acuticollis tzaidamica subsp. n. typ. ABogačev det.” [hanwritten] . Paralectotypes, 4 specimens ( ZIN) with the same first label. Bogatchev (1951) included five specimens in the type series.
Diagnosis. Body slender, shiny, brown or black, dorsally covered with short recumbent setae entirely ( Fig. 5A, B, D View FIGURE 5 ) or partially ( Figs 7A, B View FIGURE 7 ). Eyes separated by lateral margin of genae only to their midlength; narrowest portion consists of 2–3 ommatidia in width ( Fig. 3A View FIGURE 3 ). Lateral outline of head with sharp emargination between gena and epistoma ( Fig. 5E View FIGURE 5 ). Pronotum with acute, projected antero- and posterolateral angles with pointed tip; basal margin slightly serrate, not margined ( Figs 5E View FIGURE 5 , 7A View FIGURE 7 ); disc without transverse impressions along base at each side. Prothoracic hypomera finely densely wrinkled or without wrinkles, with small sparse granules, not flattened laterally ( Fig. 5C View FIGURE 5 ). Elytral interstriae granulated et least basally, each granule with short recumbent seta; rest surface of elytra can be punctured or interstriae with very small sparse tubercles. Protibiae narrow, curved, with deeply emarginated apical margin ( Fig. 5F View FIGURE 5 , 7A, B View FIGURE 7 ). Parameres narrow, acute at apex, almost straight, basal piece approximately 1.7 times as long as parameres ( Fig. 6A View FIGURE 6 ). Spiculum gastrale with oval blades ( Fig. 6C View FIGURE 6 ). Spiculum ventrale with thick stem and very short setation ( Fig. 6D View FIGURE 6 ). Ovipositor with strongly elongate and acute lobes of coxite IV, paraproct large, campanulate, with rouded walls; coxite almost completely nested in paraproct in folded position ( Figs 6E, F View FIGURE 6 ).
Composition. Penthicus acuticollis ( Reitter, 1887) , Penthicus tzaidamicus ( Bogatchev, 1951) . Formally Myladion includes 46 species and subspecies listed in the catalogue ( Iwan et al. 2020) (except for P. pavlovskyi (Reichardt, 1935) and P. truncatus Reitter, 1896 ), P. belutschistanicus ( Kaszab, 1961) , P. pakistanicus sp. nov. and Penthicus sp. from Upper Dir ( Pakistan).
Distribution. Central Asia from Southern Siberia and Eastern Kazakhstan to China and Mongolia, Pakistan.
Notes. Central Asian species of the subgenus (from eastern Kazakhstan to Mongolia and N China) have usually robust body and eyes divided by more than half length; dorsal and ventral portions of eyes are connected by very narrow bridge of one line of ommatidiae. Bogatchev (1972) believed that this subgenus is polyphyletic and only two species ( P. acuticollis and P. tzaidamicus ) should remain in Myladion s. str., and the rest should be revised and included to other subgenera.
Subgenus Discotus
Type species Opatrum dilectans Faldermann, 1836 , subsequent designation (Medvedev 1990).
Diagnosis. Basal margin of pronotum not serrated. Pronotum with widely flattened and raised lateral margins which are granulated or coarsely wrinkled. Hind wings present, species able to fly. Parameres narrow, acute and subulate in apical two thirds. Ovipositor with moderately widened rounded lobes of coxite IV and well-expressed gonostyli.
Composition. Five species, listed in the catalogue ( Iwan et al. 2020).
Distribution. From North Africa and the Afrotropic (Sahel) to Mongolia and North China.
Subgenus Penthicus
Diagnosis. Pronotum with basal margin serrate laterally. Parameres slightly shorter than or same long as basal piece of aedeagus, almost straight, strongly narrowed in apical half. Ovipositor with triangularly pointed apical lobes of coxite IV, gonostyli very small granular or rudimentary, some species ( P. pinguis , P. germanni sp. nov.) have not triangular but narrow, apically rounded lobes of coxite IV.
Composition: At least P. pinguis (= P. auliensis ) with five subspecies, P. kirgizicus (Bogatchev, 1961) , P. rufescens (Mulsant & Rey, 1859) with two subspecies, P. samgarensis Reitter, 1896 , P. oblongopunctatus ( Reitter, 1904) , P. rugosicollis ( Bogatchev, 1951) , P. zhenzhuristi ( Bogatchev, 1951) with two subspecies, P. mesopotamicus sp. nov. and P. germanni sp. nov.
Distribution. From Levant region through Southern Anatolia, Transcaucasia and plains of Central Asia to N Tian-Shan, Afghanistan, Pakistan.
Subgenus Stonavus Reitter, 1904 , subgen. resurr.
Type species Penthicus alaensis Reitter, 1896 , by subsequent designation ( Iwan & Löbl 2007)
Diagnosis. Eyes entirely divided into dorsal and ventral halves, except for some species and specimens of the corpulentus species-group. Base of pronotum not serrate. Humeral angles slightly projected or not. Parameres usually much or, rarely, slightly shorter than basal piece of aedeagus, strongly or moderately widened ( Fig. 8 View FIGURE 8 ). Ovipositor with wide or moderately rounded coxite lobes IV and with well expressed gonostyli.
Notes. We resurrect this subgenus which distinctly differs from the nomynotypical one in the not serrated base of pronotum, the shape of the ovipositor and parameres. Other seven species included in the subgenus Penthicus ( Iwan et al. 2020) must be revised: P. granuliferus ( Reichardt, 1936) , P. granulosus Ménétriés, 1849 , P. lenczyi (Kaszab, 1968) , P. lindbergi ( Kaszab, 1970) , P. netuschili ( Reitter, 1904) , P. winkleri Schuster, 1915 . We do not include them in any subgenus until a revision.
1. remotus species-group. Base of pronotum entirely margined; lateral margins of pronotum not flattened or, rarely, very narrowly flattened; elytra at apex and lateral margins of pronotum without visible setae; aedeagus moderately widened, parameres moderately narrow, slightly or strongly S-shaped ( Figs 7A–E View FIGURE 7 , 51A View FIGURE 51 , 53A View FIGURE 53 , 58A View FIGURE 58 , 64A View FIGURE 64 , 68A View FIGURE 68 ).
Composition: At least P. adustus ( Bogatchev, 1951) , P. alaensis Reitter, 1896 , P. bam sp. nov., P. bogatshevi ( Kaszab, 1970) , P. captiosus sp. nov., P. curtus ( Kaszab, 1970) , P. grabantae sp. nov., P. iranensis sp. nov., P. khorasanicus sp. nov., P. kermanicus sp. nov., P. medvedevorum (Bogatchev, 1970) , P. pazukii sp. nov., P. remotus Reitter, 1896 with two subspecies, P. telnovi sp. nov., P. zarudnyi sp. nov.
2. corpulentus species-group. Body robust, usually large (length up to 18.5 mm). Dorsal and ventral portions of eyes connected by very narrow bridge of one ommatidia width or one isolated ommatidium can be between dorsal and ventral halves of eye, or eyes entirely narrowly divided by connected gena and temple; sometimes left eye with bridge of ommatidia, and right eye entirely divided (or vice versa). Pronotum with obtuse, not projected antero- and posterolateral angles with narrowly or widely rounded tip; base entirely margined; lateral sides of pronotum widely obliquely or rarely horizontally flattened (only P. farsistanus has not flattened lateral sides of pronotum); disc slightly transversely impressed along base at each side; basal edge of pronotum not serrated. Elytra at apex and sometimes on lateral margins of pronotum with very short but distinct setae; aedeagus strongly widened ar median portion, parameres strongly narrowed from base towards apex, straight laterally ( Figs 8F–H View FIGURE 8 ).
Composition: P. corpulentus Reitter, 1896 , P. farsistanus sp. nov., P. persicus ( Schuster, 1919) , P. shokhini sp. nov., P. villiersi ( Kaszab, 1969) .
3. iners species-group. Base of pronotum medially not margined (existing margin can be only shortly interruptred at midllength; sides of pronotum flattened or not; elytra at apex and lateral margins of pronotum without visible setae; parameres wide, subparallel in basal ¾ and obliquely truncate at apex ( Figs 8I–O View FIGURE 8 ).
Composition: P. achaemenus sp. nov., P. ardoini ( Kaszab, 1969) , P. fartilis Reitter, 1899 , P. flatilatus sp. nov., P. freyi ( Schuster, 1937) , P. horni ( Schuster, 1922) , P. iners ( Ménétriés, 1832) , P. kasatkini sp. nov., P. similis ( Schuster, 1937) , P. turcomanicus ( Medvedev, 1964) .
Distribution. SE Anatolia, Iraq, Transcaucasia, Iran, Turkmenistan, Afghanistan, Tajikistan, Uzbekistan (N slopes of Alay Mts.), Western China.
Subgenus Allomyladion Bogatchev, 1972
Type species Penthicus poricollis Reitter, 1896 , original designation ( Lobothorax poricollis in the original description of the subgenus).
Notes. Species of this subgenus characterized by slender body with coarse sculpture of dorsal surface ( Bogatchev 1972).
Composition. Bogatchev (1972) included the following similar species in this subgenus, which were listed in the differential diagnosis of P. kaszabi Bogatchev, 1972 : P. poricollis Reitter, 1896 , P. zoltani G.S. Medvedev & Iwan, 2006 (= P. kaszabi , nom preocc.), P. pavlovskyi (Reichardt, 1935) and P. truncatus Reitter, 1896 . The two latter were erroneously placed in the subgenus Myladion in the catalogue ( Iwan et al. 2020).
Distribution. Southern Uzbekistan, Tajikistan and Afghanistan.
Subgenus Aulonolcus Reitter, 1904
Type species Pedinus altaicus Gebler, 1830 , by monotypy.
Notes. A monotypical subgenus, characterized by not margined lateral edges of pronotum and deep, groove-like impression along pronotal base.
Distribution. Southern Siberia, Mongolia, North China (Xinjiang, Inner Mongolia).
Subgenus Pseudopenthicinus Bogatchev, 1972
Type species Penthicus medvedev Bogatchev, 1972 , original designation.
Notes. A monotypical subgenus, characterized by almost straight base of pronotum and incompletely divided eyes.
Distribution. South Tajikistan.
Species from Iraq, Iran and Pakistan
Subgenus Penthicus Faldermann, 1836
| ZIN |
Russian Academy of Sciences, Zoological Institute, Zoological Museum |
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